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Archie Carr Center for Sea Turtle Research

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In Press

Price, J.T., J.B. Pfaller, H.B. Vander Zanden, K.L. Williams, A.B. Bolten, and K.A. Bjorndal. In press. Foraging area, not trophic position, linked to head size variation in adult female loggerhead turtles. Journal of Zoology.

Stacy N.I., K. Barnhart, and M.M. Fry.  In press.  Comparative hematology. In: Wintrobe’s Clinical Hematology.


Bjorndal, K.A., A.B. Bolten, M. Chaloupka, V.S. Saba, C. Bellini, M.A.G. Marcovaldi, A.J.B. Santos, L.F. Wurdig Bortolon, A.B. Meylan, P.A. Meylan, J. Gray, R. Hardy, B. Brost, M. Bresette, J.C. Gorham, S. Connett, B. Van Sciver Crouchley, M. Dawson, D. Hayes, C.E. Diez, R.P. van Dam, S. Willis, M. Nava, K.M. Hart, M.S. Cherkiss, A.G. Crowder, C. Pollock, Z. Hillis-Starr, F.A. Muñoz Tenería, R. Herrera-Pavón, V. Labrada-Martagón, A. Lorences, A. Negrete-Philippe, M.M. Lamont, A.M. Foley, R. Bailey, R.R. Carthy, R. Scarpino, E. McMichael, J.A. Provancha, A. Brooks, A. Jardim, M. López-Mendilaharsu, D. González-Paredes, A. Estrades, A. Fallabrino, G. Martínez-Souza, G.M. Vélez-Rubio, R.H. Boulon, Jr., J.A. Collazo, R. Wershoven, V. Guzmán Hernández, T.B. Stringell, A. Sanghera, P.B. Richardson, A.C. Broderick, Q. Phillips, M. Calosso, J.A.B. Claydon, T.L. Metz, A.L. Gordon, A.M. Landry, Jr., D.J. Shaver, J. Blumenthal, L. Collyer, B.J. Godley, A. McGowan, M.J. Witt, C.L. Campbell, C.J. Lagueux, T.L. Bethel and L. Kenyon. 2017. Ecological regime shift drives declining growth rates of sea turtles throughout the West Atlantic. Global Change Biology.  Article DOI: 10.1111/gcb.13712.  Bjorndal_et_al_GCB_2017.pdf

Lagueux, C.J., C.L. Campbell, and S. Strindberg. 2017. Artisanal green turtle (Chelonia mydas) fishery of Caribbean Nicaragua: II. Characterization and trends in size, sex, and maturity status of turtles killed, 1994-2011. Marine Biology 164:60. DOI: 10.1007/s00227-017-3092-2.

Soares L.S, A.B. Bolten, M.L. Wayne, S.T. Vilaca, F.R. Santos, M.A. dei Marcovaldi, and K.A. Bjorndal. 2017. Comparison of reproductive output of hybrid sea turtles and parental species. Marine Biology 164:9


Álvarez-Varas, R.; Berzins, R.; Bilo, K.; Chevalier, J.; Chevallier, D.; De Thoisy, B.; Fallabrino, A.; Cruz, M.G.; Kelez, S.; Lopez-Mendilaharsu, M.; Marcovaldi, M.A.; Mast, R.B.; Medrano, C.; Miranda, C.; Nalovic, M.A.; Prosdocimi, L.; Rguez-Barón, J.M.; Santos, A.; Soares, L.S.; Thome, J.; Vallejo, F.; Vélez-Rubio, G. 2016. Sea turtles of South America. In: SWoT Report XI. The State of the World’s Sea Turtles. pp 14-27.

Bjorndal, K.A., M. Chaloupka, V.S. Saba, C.E. Diez, R.P. van Dam, B.H. Krueger, J.A. Horrocks, A.J.B. Santos, C. Bellini, M.A.G. Marcovaldi, M. Nava, S. Willis, B.J. Godley, S. Gore, L.A. Hawkes, A. McGowan, M.J. Witt, T.B. Stringell, A. Sanghera, P.B. Richardson, A.C. Broderick, Q. Phillips, M.C. Calosso, J.A.B. Claydon, J. Blumenthal, F. Moncada, G. Nodarse, Y. Medina, S.G. Dunbar, L.D. Wood, C.J. Lagueux, C.L. Campbell, A.B. Meylan, P.A. Meylan, V.R. Burns Perez, R.A. Coleman, S. Strindberg, V. Guzmán-H., K.M. Hart, M.S. Cherkiss, Z. Hillis-Starr, I.F. Lundgren, R.H. Boulon, Jr., S. Connett, M.E. Outerbridge, and A.B. Bolten.  2016.  Somatic growth dynamics of West Atlantic hawksbill sea turtles: a spatio-temporal perspective.  Ecosphere 7(5):e01279.  DOI: 10.1002/ecs2.1279.  Bjorndal_et_al_Ecosphere_2016.pdf

Gil MA, Pfaller JB. 2016. Oceanic barnacles act as foundation species on plastic debris: implications for marine dispersal. Scientific Reports 6:19987

Huertas, V. and C.J. Lagueux. 2016. First recorded mass stranding of the short-finned pilot whale (Globicephala macrorhynchus) on the Caribbean coast of Nicaragua. Aquatic Mammals 42:27-34. doi: 10.1578/AM.42.1.2016.27

Kaplan, Amy J., Nicole I. Stacy, Elliott Jacobson, Carolina R. Le-Bert, Hendrik H. Nollens, Francesco C. Origgi, Linda G. Green, Shadi Bootorabi, Alan Bolten, Jorge A. Hernandez. 2016.  Development and validation of a competitive enzyme-linked. immunosorbent assay for the measurement of total plasma immunoglobulins in healthy loggerhead sea (Caretta caretta) and green turtles (Chelonia mydas).  Journal of Veterinary Diagnostic Investigation 28:5-11.

Lagueux, C.J. and C.L. Campbell. 2016. Caught in a net: Green turtles and the turtle people of Nicaragua. The State of the World’s Sea Turtles (SWOT) Report 11:34-35.

Mariela Pajuelo, Karen A. Bjorndal, Michael D. Arendt, Allen M. Foley, Barbara A. Schroeder, Blair E. Witherington, Alan B. Bolten. 2016.  Long‑term resource use and foraging specialization in male loggerhead turtles.  Mar Biol 163:235 pajuelo-et-al-2016.pdf

Rees, A.F.,  J. Alfaro-Shigueto, P.C.R. Barata, K.A. Bjorndal, A.B. Bolten, J. Bourjea, A.C. Broderick, L.M. Campbell, L. Cardona, C. Carreras, P. Casale, S.A. Ceriani, P.H. Dutton, T. Eguchi, A. Formia, M.M.P.B. Fuentes, W.J. Fuller, M. Girondot, M.H. Godfrey, M. Hamann, K.M. Hart, G.C. Hays, S. Hochscheid, Y. Kaska, M.P. Jensen, J.C. Mangel, J.A. Mortimer, E. Naro-Maciel, C.K.Y. Ng, W.J. Nichols, A.D. Phillott, R.D. Reina, O. Revuelta, G. Schofield, J.A. Seminoff, K. Shanker, J. Tomás, J.P. van de Merwe, K.S. Van Houtan, H.B. Vander Zanden, B.P. Wallace, K.R. Wedemeyer-Strombel, T.M. Work, and B.J. Godley.  2016.  Are we working towards global research priorities for management and conservation of sea turtles?  Endangered Species Research 31: 337–382.

Strindberg, S., R.A. Coleman, V.R. Burns Perez, C.L. Campbell, I. Majil, and J. Gibson. 2016. In-water assessments of sea turtles at Glover’s Reef Atoll, Belize. Endangered Species Research 31: 211-225. doi: 10.3354/esr00765.

Hannah B. Vander Zanden, Alan B. Bolten, Anton D. Tucker, Kristen M. Hart, Margaret M. Lamont, Ikuko Fujisaki, Kimberly J. Reich, David S. Addison, Katherine L. Mansfield, Katrina F. Phillips, Mariela Pajuelo, and Karen A. Bjorndal. 2016.  Biomarkers reveal sea turtles remained in oiled areas following the Deepwater Horizon oil spill.  Ecological Applications, 26(7), 2016, pp. 2145–2155  Vander_Zanden_et_al_2016_Ecol_Appl.pdf


Avens, L., L.R. Goshe, L. Coggins, M.L. Snover, M. Pajuelo, K.A. Bjorndal, and A.B. Bolten.  2015. Age and size at maturation and adult stage duration for loggerhead sea turtles in the western North Atlantic. Marine Biology 162:1749-1767.

Hammerschlag, N., A.C. Broderick, J.W. Coker, M.S. Coyne, M. Dodd, M.G. Frick, M.H. Godfrey, B.J. Godley, D.B. Griffin, K. Hartog, S.R. Murphy, T.M. Murphy, E.R. Nelson, K.L. Williams, M.J. Witt, and L.A. Hawkes.  2015. Evaluating the landscape of fear between apex predatory sharks and mobile sea turtles across a large dynamic seascape. Ecology 96:2117–2126.

Rocha Júnior JC, Pfaller JB, Corbetta R, Veríssimo L.  2015.  Parasitic isopods associated with sea turtles nesting in Brazil. Journal of the Marine Biology Association of the U.K. 95(5):973-981

Shamblin, B.M., P.H. Dutton, K.A. Bjorndal, A.B. Bolten, E. Naro-Maciel, A.J.B. Santos, C. Bellini, C. Baptistotte, M.Â. Marcovaldi, and C.J. Nairn. 2015. Deeper mitochondrial sequencing reveals cryptic diversity and structure in Brazilian green turtle rookeries. Chelonian Conservation and Biology 14:167-172.

Thomson, J.A., A. Gulick, M. R. Heithaus. 2015. Intraspecific behavioral dynamics in a green turtle Chelonia mydas foraging aggregation. Marine Ecology Progress Series 532: 243–256.

Vander Zanden, H.B., A.D. Tucker, M. Hart, M.M. Lamont, I. Fujisaki, D.S. Addison, K.L. Mansfield, K.F. Phillips, M.B. Wunder, G.J. Bowen, M. Pajuelo, A.B. Bolten, and K.A. Bjorndal. 2015. Determining origin in a migratory marine vertebrate: a novel method to integrate stable isotopes and satellite tracking. Ecological Applications 25:320–335.  Vander_Zanden_et_al_Ecol_Appl_2015.pdf

Zárate, P.M., K.A. Bjorndal, J.A. Seminoff, P.H. Dutton, and A.B. Bolten. 2015. Somatic growth rates of green turtles and hawksbills in the Galápagos Islands. Journal of Herpetology 49:641–648.


Bjorndal, K.A., J. Parsons, W. Mustin, and A.B. Bolten. 2014. Variation in age and size at sexual maturity in Kemp’s ridley sea turtles. Endangered Species Research 25:57–67.

Bolten, A.B. and K.A. Bjorndal. 2014.  Sea turtles in Bahamian waters.  Pages 210-213 in G.C. Ray and J. McCormick-Ray (editors), Marine Conservation: Science, Policy, and Management.  Blackwell Scientific, London.

Campbell, CL. 2014. Conservation status of hawksbill turtles in the Wider Caribbean, Western Atlantic, and Eastern Pacific regions. IAC Secretariat Pro Tempore, Virginia USA. 76p. English: Spanish:

Kaufman, T.J., M. Pajuelo, K.A. Bjorndal, A.B. Bolten, J.B. Pfaller, K.L. Williams, and H.B. Vander Zanden. 2014. Mother-egg stable isotope conversion factors and effects of lipid extraction and ethanol preservation on loggerhead eggs. Conservation Physiology 2: doi:10.1093/conphys/cou049.

Lagueux, C.J., C.L. Campbell, and S. Strindberg. 2014. Artisanal green turtle, Chelonia mydas, fishery of Caribbean Nicaragua: I. Catch rates and trends, 1991-2011. PLoS ONE 9(4): e94667. doi:10.1371/journal.pone.0094667.

López-Castro, M.C., K.A. Bjorndal, and A.B. Bolten. 2014. Evaluation of scute thickness to infer life history records in the carapace of green and loggerhead turtles. Endangered Species Research 24:191-196.

López-Castro, M.C., K.A. Bjorndal, G.D. Kamenov, and A.B. Bolten. 2014. Identifying oceanic foraging grounds of sea turtles in the Atlantic using lead isotopes. Marine Biology 161:2269–2278.

Marcovaldi, M.A.G. dei, A.J.B. Santos, A.S. Santos, L.S. Soares, G.G. Lopez, M.H. Godfrey, M. López-Mendilaharsu, and M.M.P.B. Fuentes. 2014. Spatio-temporal variation in the incubation duration and sex ratio of hawksbill hatchlings: Implication for future management. Journal of Thermal Biology 44:70–77.

Naro‐Maciel, E., B.N. Reid, S.E. Alter, G. Amato, K.A. Bjorndal, A.B. Bolten, M. Martin, C.J. Nairn, B. Shamblin, and O. Pineda-Catalan. 2014. From refugia to rookeries: phylogeography of Atlantic green turtles. Journal of Experimental Marine Biology and Ecology 461:306–316.

Pfaller, J.B., J. Alfaro-Shigueto, B. Giffoni, T. Ishihara, J.C. Mangel, S.H. Peckham, K.A. Bjorndal, and J.A. Baeza. 2014. Social monogamy in the crab Planes major, a facultative symbiont of loggerhead sea turtles. Journal of Experimental Marine Biology and Ecology 461:124–132.

Pfaller, J.B., J. Alfaro-Shigueto, G.H. Balazs, T. Ishihara, K. Kopitsky, J.C. Mangel, S.H. Peckham, A.B. Bolten, and K.A. Bjorndal. 2014. Hitchhikers reveal cryptic host behavior: new insights from the association between Planes major and sea turtles in the Pacific Ocean. Marine Biology 161:2167–2178.

Roark A.M. and K.A. Bjorndal. 2014. Bridging developmental boundaries: lifelong dietary patterns modulate life histories in a parthenogenetic insect. PLoS ONE 9(11): e111654. doi: 10.1371/journal.pone.0111654.

Shamblin, BM, AB Bolten, FA Abreu-Grobois, KA Bjorndal, L Cardona, C Carreras, M Clusa, C Monzon-Arguello, CJ Nairn, JT Nielsen, R Nel, LS Soares, KR Stewart, ST Vilaca, O Turkozan, C Yilmaz, PH Dutton.  2014.  Geographic patterns of genetic variation in a broadly distributed marine vertebrate: New insights into loggerhead turtle stock structure from expanded mitochondrial DNA sequences.  PLoS ONE 9(1): e85956. doi:10.1371/journal.pone.0085956.  Shamblin_etal_PLOSOne_2014.pdf (supplemental tables Shamblin_etal_PLOSOne_2014_Suppl_Tables)

Vandeperre, F., A. Aires-da-Silva, M. Santos, R. Ferreira, A.B. Bolten, R.S. Santos, and P. Afonso. 2014. Demography and ecology of blue shark (Prionace glauca) in the central North Atlantic. Fisheries Research 153:89–102.

Vander Zanden, H.B., J.B. Pfaller, K.J. Reich, M. Pajuelo, A.B. Bolten, K.L. Williams, M.G. Frick, B.M. Shamblin, C.J. Nairn, and K.A. Bjorndal.  2014.  Foraging areas differentially affect reproductive output and interpretation of trends in abundance of loggerhead turtles.  Marine Biology 161:585-598.  Vander_Zanden_et_al_2014_MarBiol.pdf

Vander Zanden, H.B., A.D. Tucker, A.B. Bolten, K.J. Reich, and K.A. Bjorndal. 2014. Stable isotopic comparison between loggerhead sea turtle tissues. Rapid Communications in Mass Spectrometry 28:2059–2064.

Williams, N.C., K.A. Bjorndal, M.M. Lamont, and R.R. Carthy. 2014. Winter diets of immature green turtles (Chelonia mydas) on a northern feeding ground: integrating stomach contents and stable isotope analyses. Estuaries and Coasts 37:986–994.

Zardus, J.D., D.T. Lake, M.G. Frick and P.D. Rawson. 2014. Deconstructing an assemblage of “turtle” barnacles: species assignments and fickle fidelity in Chelonibia. Marine Biology 2014.  Marine Biology 161:45-59.   Zardus_etal_2014.pdf


Avens, L., L.R. Goshe, M. Pajuelo, K.A. Bjorndal, B.D. MacDonald, G.E. Lemons, A.B. Bolten, and J.A. Seminoff.  2013.  Complementary skeletochronology and stable isotope analyses offer new insight into juvenile loggerhead sea turtle (Caretta caretta) oceanic stage duration and growth dynamics.  Marine Ecology Progress Series 491: 235–251.

Bjorndal, K.A., J. Parsons, W. Mustin, and A.B. Bolten. 2013. Threshold to maturity in a long-lived reptile: interactions of age, size and growth. Marine Biology 160:607-616. Bjorndal_et_al_MarBiol_2013.pdf

Bjorndal, K.A., B.A. Schroeder, A.M. Foley, B.E. Witherington, M. Bresette, D. Clark, R.M. Herren, M.D. Arendt, J.R. Schmid, A.B. Meylan, P.A. Meylan, J.A. Provancha, K.M. Hart, M.M. Lamont, R.R. Carthy, and A.B. Bolten.  2013.  Temporal, spatial, and body size effects on growth rates of loggerhead sea turtles (Caretta caretta) in the Northwest Atlantic.  Marine Biology 160:2711-2721.  Bjorndal_et_al_MarBiol_2013b.pdf

Frick, M.G. 2013. A rejoinder and addendum to Hayashi (2011) regarding the systematics and biology of the turtle and whale barnacles (Cirripedia: Balanomorpha: Coronuloidea). Journal of the Marine Biological Association of the United Kingdom 93:183-187. Frick_2012.pdf

Frick, M.G.  2013.  Clarification on the nomenclatural availability of the turtle barnacle genus Chelolepas (Cirripedia: Balanomorpha: Coronuloidea), a corrigendum to Hayashi (2012).  Journal of the Marine Biological Association of the United Kingdom 93:1579-1580. Frick-2013.pdf

Frick, M.G. and J.B. Pfaller. 2013.  Sea turtle epibiosis. Pages 399-426 in The Biology of Sea Turtles Volume III. CRC Press.

Griffin, D.B.,  S.R. Murphy, M.G. Frick, A.C. Broderick , J.W. Coker, M.S. Coyne, M.G. Dodd, M.H. Godfrey, B.J. Godley, L.A. Hawkes, T.M. Murphy, K.L. Williams and M.J. Witt. 2013. Foraging habitats and migration corridors utilized by a recovering subpopulation of adult female loggerhead sea turtles: implications for conservation. Marine Biology 160:3071-3086.  Griffin_etal_2013.pdf

López-Castro, M.C., K.A. Bjorndal, G.D. Kamenov, R. Zenil-Ferguson, and A.B. Bolten. 2013. Sea turtle population structure and connections between oceanic and neritic foraging areas in the Atlantic revealed through trace elements. Marine Ecology Progress Series 490: 233–246.

Pfaller, J.B., K.A. Bjorndal, M. Chaloupka, K.L. Williams, M.G. Frick, and A.B. Bolten.  2013.  Accounting for imperfect detection is critical for inferring marine turtle nesting population trends.  PLoS ONE 8(4): e62326. doi:10.1371/journal.pone.0062326  Pfaller-et-al.-2013-PLOS-ONE.pdf

Rousselet, E., N.I. Stacy, K. LaVictoire, B.M. Higgins, M.E. Tocidlowski, J. Flanagan, and C.A.J. Godard-Codding. 2013. Hematology and plasma biochemistry analytes in five age groups of immature, captive-reared loggerhead sea turtles (Caretta caretta). Journal of Zoo and Wildlife Medicine 44:859-874.

Stacy, N.I., C.J. Innis, and J.A. Hernandez. 2013.  Development and evaluation of three mortality prediction indices for cold-stunned Kemp’s ridley sea turtles (Lepidochelys kempii).  Conservation Physiology 1: doi: 10.1093/conphys/cot003.

Vander Zanden, H.B., K.E. Arthur, A.B. Bolten, B.N. Popp, C.J. Lagueux, E. Harrison, C.L. Campbell, and K.A. Bjorndal. 2013. Trophic ecology of a green turtle breeding population. Marine Ecology Progress Series 476:237-249. Vander_Zanden_et_al_2013_MEPS.pdf

Vander Zanden, H.B., K.A. Bjorndal, and A.B. Bolten.  2013.  Temporal consistency and individual specialization in resource use by green turtles in successive life stages.  Oecologia 173:767-777. HVZ_Oecologia_2013.pdf.

Vilaca S.T., P. Lara-Ruiz, M.A. Marcovaldi, L.S. Soares, and F.R. Santos.  2013.  Population origin and historical demography in hawksbill (Eretmochelys imbricata) feeding and nesting aggregates from Brazil. Journal of Experimental Marine Biology and Ecology 446:334–344.

Zárate, P., K.A. Bjorndal, M. Parra, P.H. Dutton, J.A. Seminoff, and A.B. Bolten.  2013. Hatching and emergence success of green turtle (Chelonia mydas) nests in the Galápagos Islands.  Aquatic Biology 19:217-229.


Avens L.A., L. Goshe, C. Harms, E.T. Anderson, A.G. Hall, W.M. Cluse, M.H. Godfrey, J. Braun-McNeill, B. Stacy, R. Bailey, and M.M. Lamont. 2012. Population characteristics, age structure, and growth dynamics of neritic juvenile green turtles in the northeastern Gulf of Mexico Marine Ecology Progress Series 458:213-229. Avens_et_al_2012_MEPS.pdf

Carthy, R.R.  2012.  The leatherbacks of Grande Riviere, Trinidad.  Florida Environmental Outreach 3:10-13.

Frankel, N.S., H.B. Vander Zanden, K.J. Reich, K.L. Williams, and K.A. Bjorndal. 2012. Mother-offspring stable isotope discrimination in loggerhead sea turtles Caretta caretta. Endangered Species Research 17:133–138. Frankel_etal_ESR_2012.pdf

Hart, K., M.M. Lamont, I. Fujisaki, A.D. Tucker, and R.R. Carthy. 2012. Common coastal foraging areas for loggerheads in the Gulf of Mexico: opportunities for marine conservation. Biological Conservation 145:185-194. Hart_et_al_2012.pdf

Lamont, M.M., R.R. Carthy and I. Fujisaki. 2012. Declining reproductive parameters highlight conservation needs for loggerhead turtles (Caretta caretta) in the northern Gulf of Mexico. Chelonian Conservation and Biology  11(2):190-196.  Lamont_et_al_2012_CCB.pdf

Pajuelo, M., K.A. Bjorndal, K.J. Reich, M.D. Arendt, and A.B. Bolten. 2012. Distribution of foraging habitats of male loggerhead turtles (Caretta caretta) as revealed by stable isotopes and satellite telemetry. Marine Biology 159:1255–1267. Pajuelo_et_al_MarBiol_2012.pdf

Pajuelo, M., K. A. Bjorndal, K. J. Reich, H. B. Vander Zanden, L. A. Hawkes, and A. B. Bolten. 2012. Assignment of nesting loggerhead turtles to their foraging areas in the Northwest Atlantic using stable isotopes. Ecosphere 3(10):89. Pajuelo_etal_Ecosphere_2012.pdf

Salmon, M., R. Carthy, C.M.F. Lohmann, K.J. Lohmann and J. Wyneken. 2012. Collecting a sample of loggerhead sea turtle hatchlings before a natural emergence does not reduce nest productivity. Endangered Species Research 16:295-299.

Shamblin, B.M., K.A. Bjorndal, A.B. Bolten, Z.M. Hillis-Starr, I. Lundgren, E. Naro-Maciel, and C.J. Nairn. 2012. Mitogenomic sequences better resolve stock structure of southern Greater Caribbean green turtle rookeries. Molecular Ecology 21:2330-2340. Shamblin_etal_MEC_2012.pdf

Shamblin, B.M., K.A. Bjorndal, A.B. Bolten, and C.J. Nairn. 2012. Natal homing by an adult male green turtle at Tortuguero, Costa Rica. Marine Turtle Newsletter 134:21-22. Shamblin_etal_MTN_2012.pdf

Shamblin, B.M., A.B. Bolten, K.A. Bjorndal, P.H. Dutton, J.T. Nielsen, F.A. Abreu-Grobois, K.J. Reich, B.E. Witherington, D.A. Bagley, L.M. Ehrhart, A.D. Tucker,D.S. Addison, A.Arenas, C. Johnson, R.R. Carthy, M.M. Lamont, M.G. Dodd, M.S. Gaines, E.LaCasella, C.J. Nairn. 2012. Expanded mitochondrial control region sequences increase resolution of stock structure among North Atlantic loggerhead turtlerookeries Marine Ecology Progress Series 469:145-160. Shamblin_et_al_MEPS_2012.pdf

Vander Zanden, H.B, K.A. Bjorndal, W. Mustin, J.M. Ponciano, and A.B. Bolten. 2012. Inherent variation in stable isotope values and discrimination factors in two life stages of green turtles. Physiological and Biochemical Zoology 85:431–441. Vander_Zanden_et_al_PBZ_2012.pdf

Vander Zanden, H.B., K.A. Bjorndal, P.W. Inglett and A.B. Bolten. 2012. Marine-derived nutrients from green turtle nests subsidize terrestrial beach ecosystems. Biotropica 44:294-301. Vander_Zanden_et_al_-2012_Biotropica.pdf


Almeida, A.D.P., L.M.P. Moreira, S.C. Bruno, J.C.A. Thomé, A.S. Martins, A.B. Bolten, and K.A. Bjorndal. 2011. Green turtle nesting on Trindade Island, Brazil: abundance, trends and biometrics. Endangered Species Research 14:193-201. Almeida_etal_ESR_2011.pdf

Bjorndal, K.A., B.W. Bowen, M. Chaloupka, L.B. Crowder, S.S. Heppell, C.M. Jones, M.E. Lutcavage, D. Policansky, A.R. Solow, and B.E. Witherington. 2011. From crisis to opportunity: Better science needed for restoration in the Gulf of Mexico. Science 331:537-538. Bjorndal_et_al_Science_2011.pdf

Bjorndal, K.A. 2011. Introduction to the 2011 Reissue of the book So Excellent a Fishe by Archie Carr. University Press of Florida, Gainesville. Pages XI-XXI.

Bolten, A.B., L.B. Crowder, M.G. Dodd, S.L. MacPherson, J.A. Musick, B.A. Schroeder, B.E. Witherington, K.J. Long, and M.L. Snover. 2011 (online 2010). Quantifying multiple threats to endangered species: an example from loggerhead sea turtles. Frontiers in Ecology and the Environment 9:295–301. Bolten_et_al_Frontiers_2011.pdf

Ferreira, R., H. Martins, A. Bolten and K. Erzini. 2011. Turtle troubles. Global Marine Environment 14:22-23.

Frick, M.G., K. Kopitsky, A.B. Bolten, K.A. Bjorndal, and H.R. Martins. 2011. Sympatry in grapsoid crabs (genera Planes and Plagusia) from olive ridley sea turtles (Lepidochelys olivacea), with descriptions of crab diets and masticatory structures. Marine Biology 158:1699–1708. Frick et al MarBiol 2011.pdf

Peterson, C.H., F.C. Coleman, J.B.C. Jackson, R.E. Turner, G.T. Rowe, R.T. Barber, K.A. Bjorndal, R.S. Carney, R.K. Cowen, J.M. Hoekstra, J.T. Hollibaugh, S.B. Laska, R.A. Luettich, Jr., C.W. Osenberg, S.E. Roady, S. Senner, J.M. Teal, and P. Wang. 2011. A once and future Gulf of Mexico ecosystem: Restoration recommendations of an expert working group. Pew Environment Group, Washington DC. 112 pp. Petersonetal-GOM-report_2011.pdf

Ross, A. and M.G. Frick. 2011. Nomenclatural emendations of the family-group names Cylindrolepadinae, Stomatolepadinae, Chelolepadinae, Cryptolepadinae, and Tubicinellinae of Ross and Frick, 2007—including current definitions of family-groups within the Coronuloidea (Cirripedia: Balanomorpha). Zootaxa 3106: 60-66.

Shamblin, B.M., M.G. Dodd, D.A. Bagley, L.M. Ehrhart, A.D. Tucker, C. Johnson, R.R. Carthy, R.A. Scarpino, E. McMichael, D.S. Addison, K.L. Williams, M.G. Frick, S. Ouellette, A.B. Meylan, M.H. Godfrey, S.R. Murphy, and C.J. Nairn.  2011.  Genetic structure of the southeastern United States loggerhead turtle nesting aggregation: evidence of additional structure within the peninsular Florida recovery unit.  Marine Biology 158:571–587.

Stacy, N.I., A.R. Alleman, and K. Sayler.  2011.  Diagnostic hematology of reptiles.  Clinics in Laboratory Medicine 31:87-108.

Wallace, B.P., A.D. DiMatteo, A.B. Bolten, M.Y. Chaloupka, B.J. Hutchinson, F. A. Abreu-Grobois, J.A. Mortimer, J.A. Seminoff, D. Amorocho, K.A. Bjorndal, J. Bourjea, B.W. Bowen, R. Briseño Dueñas, P. Casale, B.C. Choudhury, A. Costa, P.H. Dutton, A. Fallabrino, E.M. Finkbeiner, A. Girard, M. Girondot, M. Hamann, B.J. Hurley, M. López-Mendilaharsu, M.A. Marcovaldi, J.A. Musick, R. Nel, N.J. Pilcher, S. Troëng, B. Witherington, and R.B. Mast. 2011. Global conservation priorities for marine turtles. PLoS ONE10.1371/journal.pone.0024510. Wallace_et_al_PLoSONE_2011.pdf


Atkins, A., E. Jacobson, J. Hernandez, A. Bolten, and X. Lu. 2010. Assessment of a portable point-of-care (VETSCAN VS20) biochemical analyzer for measuring plasma biochemical levels in free-living loggerhead sea turtles (Caretta caretta). Journal of Zoo and Wildlife Medicine 41:585-593. Atkins_et_al_JZoo&WildlMed_2010.pdf

Bjorndal, K.A., K.J. Reich, and A.B. Bolten. 2010. Effect of repeated tissue sampling on growth rates of juvenile loggerhead turtles (Caretta caretta). Diseases of Aquatic Organisms. Bjorndal_et_al_DAO_2010.pdf

Bjorndal, K.A. and A.B. Bolten. 2010. Hawksbill sea turtles in seagrass pastures: success in a peripheral habitat. Marine Biology 157:135-145. Bjorndal_&_Bolten_MarBiol_2010.pdf

Ferreira, R.L., H.R. Martins,A.B. Bolten, M.A. Santos, and K. Erzini. 2010. Influence of environmental and fishery parameters on loggerhead sea turtle bycatch in the longline fishery in the Azores Archipelago and implications for conservation. Journal of the Marine Biological Association of the United Kingdom. Published online first in 2010: doi:10.1017/S0025315410000846. Ferreira_et_al_MarBiolAssoc_2011.pdf

Hamann, M., M.H. Godfrey, J.A. Seminoff, K. Arthur, P.C.R. Barata, K.A. Bjorndal, A.B. Bolten, A.C. Broderick, L.M. Campbell, C. Carreras, P. Casale, M. Chaloupka, S.K.F. Chan, M.S. Coyne, L.B. Crowder, C.E. Diez, P.H. Dutton, S.P. Epperly, N.N. FitzSimmons, A. Formia, M. Girondot, G.C. Hays, C. I-Jiunn, Y. Kaska, R. Lewison, J.A. Mortimer, W.J. Nichols, R.D. Reina, K. Shanker, J.R. Spotila, J. Tomás, B.P. Wallace, T.M. Work, J. Zbinden, and B.J. Godley. 2010. Global research priorities for sea turtles: informing management and conservation in the 21st century. Endangered Species Research 11:245–269. Hamann_et_al_ESR_2010.pdf

National Research Council. 2010. Assessment of sea-turtle status and trends: integrating demography and abundance. National Academies Press, Washington, DC. 162 pages. Authors: K.A. Bjorndal (chair), B.W. Bowen, M. Chaloupka, L.B. Crowder, S.S. Heppell, C.M. Jones, M.E. Lutcavage, D. Policansky, A.R. Solow, and B.E. Witherington. National_Research_Council_2010.pdf

Moncada, F., F.A. Abreu-Grobois, D. Bagley, K.A. Bjorndal, A.B. Bolten, J.A. Camiñas, L. Ehrhart, A. Muhlia-Melo, G. Nodarse, B.A. Schroeder, J. Zurita, and L.A. Hawkes. 2010. Movement patterns of loggerhead turtles Caretta caretta in Cuban waters inferred from flipper tag recaptures. Endangered Species Research 11: 61–68. Moncada_et_al_ESR_2010.pdf

Osborne, A.G., E.R. Jacobson, M. Bresette, D. Singewald, R. Scarpino, and A.B. Bolten. 2010. Reference intervals and relationships between health status, carapace length, body mass, and water temperature, and concentrations of plasma protein and protein fractions of the Atlantic loggerhead sea turtle (Caretta caretta) and the green turtle (Chelonia mydas). Journal of the American Veterinary Medical Association 237:561-567. Osborne_et_al_JAVMA_2010.pdf

Pajuelo, M., K.A. Bjorndal, J. Alfaro-Shigueto, J.A. Seminoff, J.C. Mangel, and A.B. Bolten. 2010. Stable isotope variation in loggerhead turtles reveals Pacific-Atlantic oceanographic differences. Marine Ecology Progress Series 417:277-285. Pajuelo_et_al_MEPS_2010.pdf

Reich, K.J., K.A. Bjorndal, M.G. Frick, B.E. Witherington, C. Johnson, and A.B. Bolten. 2010. Polymodal foraging in adult female loggerheads (Caretta caretta). Marine Biology 157:113-121. Reich_et_al_MarBiol_2010.pdf

Reis, E.C., L.S. Soares, S.M. Vargas, F.R. Santos, R.J. Young, K.A. Bjorndal, A.B. Bolten, and G. Lôbo-Hajdu. 2010. Genetic composition, population structure and phylogeography of the loggerhead sea turtle: colonization hypothesis for the Brazilian rookeries. Conservation Genetics 11:1467–1477. Reis_et_al_ConsGen_2010.pdf

Senko, J., V. Koch, W.M. Megill, R.R. Carthy, R.P. Templeton, and W.J. Nichols.  2010.  Fine scale daily movements and habitat use of East Pacific green turtles at a shallow coastal lagoon in Baja California Sur, Mexico.  Journal of Experimental Marine Biology and Ecology 391:92-100.

Stacy, B.A., A.M. Foley, E. Greiner, L.H. Herbst, A. Bolten, P. Klein, C.A. Manire, and E.R. Jacobson.  2010.  Spirorchiidiasis in stranded loggerhead Caretta caretta and green turtles Chelonia mydas in Florida (USA): host pathology and significance.. Diseases of Aquatic Organisms 89:237-259. Stacy_et_al_DAO_2010.pdf

Stacy, B.A., T. Frankovich, E. Greiner, A.R. Alleman, L.H. Herbst, P. Klein, A. Bolten, A. McIntosh, and E.R. Jacobson. 2010. Detection of spirorchiid trematodes in gastropod tissues by polymerase chain reaction: preliminary identification of an intermediate host of Learedius learedi.Journal of Parasitology 96:752-757. Stacy_et_al_JParasitol_2010.pdf

Vander Zanden, H.B., K.A. Bjorndal, K.J. Reich, and A.B. Bolten. 2010. Individual specialists in a generalist population: results from a long-term stable isotope series. Biology Letters 6:711-714. Vander_Zanden_et_al_2010_BioLett.pdf

Wabnitz, C.C.C., G. Balazs, S. Beavers, K.A. Bjorndal, A.B. Bolten, V. Christensen, S. Hargrove, and D. Pauly. 2010. Ecosystem structure and processes at Kaloko Honokōhau, focusing on the role of herbivores, including the green sea turtle Chelonia mydas, in reef resilience. Marine Ecology Progress Series 420:27-44. Wabnitz_et_al_MEPS_2010.pdf

Wallace, B.P., A.D. DiMatteo, B.J. Hurley, E.M. Finkbeiner, A.B. Bolten, M.Y. Chaloupka, B.J. Hutchinson, F.A. Abreu-Grobois, D. Amorocho, K.A. Bjorndal, J. Bourjea, B.W. Bowen, R. Briseno Duenas, P. Casale, B.C. Choudhury, A. Costa, P.H. Dutton, A. Fallabrino, A. Girard, M. Girondot, M.H. Godfrey, M. Hamann, M. Lopez-Mendilaharsu, M.Angela. Marcovaldi, J.A. Mortimer, J.A. Musick, R. Nel, N.J. Pilcher, J.A. Seminoff, S. Troeng, B. Witherington, and R.B. Mast. 2010. Regional management units for marine turtles: a novel framework for prioritizing conservation and research across multiple scales. PLoS One 5(12): e15465:1-11. Wallace_et_al_PLoSONE_2010.pdf


Bjorndal, K.A. and A.B. Bolten. 2009. Policy changes protect sea turtles in The Bahamas: Long-term efforts rewarded. State of the World’s Turtles 5:16-17.

Frick, M.G., K.L. Williams, A.B. Bolten, K.A. Bjorndal, and H.R. Martins. 2009. Foraging ecology of oceanic-stage loggerhead turtles (Caretta caretta), opportunistic carnivores. Endangered Species Research 9:91-97. Frick_et_al_EndangSpeciesRes_2009.pdf

Herbst L.H, J. Lenz, K. Van Doorslaer, Z. Chen, B.A. Stacy, J.F Wellehan Jr, C.A. Manire, and R.D. Burk. 2009. Genomic characterization of two novel reptilian papillomaviruses, Chelonia mydas papillomavirus 1 and Caretta caretta papillomavirus 1. Virology 383(1):131-5.

Mast, R.B., B. Wallace, B.J. Hutchinson, M. Chaloupka, A.B. Bolten, and N.J. Pilcher.  2009.  IUCN-SSC Marine Turtle Specialist Group Quarterly Report: Progress from the Fifth Burning Issues Workshop (BI-5).  Marine Turtle Newsletter 126:17-19.

Roark, A.M., K.A. Bjorndal, and A.B. Bolten. 2009. Compensatory responses to food restriction in juvenile green turtles (Chelonia mydas). Ecology 90:2524–2534. Roark_et_al_Ecology_2009.pdf

Roark, A.M., C. Leeuwenburgh, K.A. Bjorndal, and A.B. Bolten. 2009. Biochemical indices as correlates of recent growth in juvenile green turtles (Chelonia mydas). Journal of Experimental Marine Biology and Ecology 376:59–67. Roark_et_al_JEMBE_2009.pdf‎

Turtle Expert Working Group. 2009. An Assessment of the Loggerhead Turtle Population in the Western North Atlantic Ocean. NOAA Technical Memorandum NMFS-SEFSC-575, 142 pages.  Turtle Expert Working Group:  N. Thompson, A. Bolten, M. Dodd, S. Epperly, A. Foley, M. Godfrey, H. Haas, S. MacPherson, K. Mansfield, P. Pritchard, P. Richards, C. Sasso, M. Snover, J. Vaughan, J. Wyneken, and J.C. Zurita.


Barrow, L.M., K.A. Bjorndal, and K.J. Reich. 2008. Effects of preservation method on stable carbon and nitrogen isotope values. Physiological and Biochemical Zoology 81:688–693. Barrow_et_al_PBZ_2008.pdf‎

Bjorndal, K.A. and A.B. Bolten. 2008. Annual variation in source contributions to a mixed stock: implications for quantifying connectivity. Molecular Ecology 17:2185–2193. Bjorndal&Bolten_MEC_2008.pdf

Bjorndal, K.A., T. Clovis, K.J. Reich, G. Alkins, P.J. Eliazar, and A.B. Bolten. 2008. Juvenile hawksbill tagged in The Bahamas nests inTobago. Marine Turtle Newsletter 122:10-11. Bjorndal_et_al_MTN_2008.pdf

Chaloupka, M., K.A. Bjorndal, G.H. Balazs, A.B. Bolten, L.M. Ehrhart, C.J. Limpus, H. Suganuma, S. Troëng, and M. Yamaguchi. 2008. Encouraging outlook for recovery of a once severely exploited marine megaherbivore. Global Ecology and Biogeography 17:297-304. Chaloupka_etal_GEB_2008.pdf

Eisenhawer, E., C.H. Courtney, R.E. Raskin, and E.R. Jacobson.  2008.  Relationship between separation-time of plasma from whole blood on plasma biochemical analytes of loggerhead sea turtles.  Journal of Zoo and Wildlife Medicine 39: 208-215.

Herbst, L.H., S. Lemaire, A.R. Ene, D.J. Heslin, L.M. Ehrhart, D.A. Bagley, P.A. Klein, and J. Lenz.  2008.  ELISA detection of plasma antibodies to baculovirus-expressed glycoprotein H of chelonid FP-associated herpesvirus and its relationship to the prevalence of fibropapillomatosis in sea turtles.  Clinical and Vaccine Immunology 5:843-851.

McMichael, E., J.A. Seminoff and R.R. Carthy.  2008.  Growth rates of wild green turtles, Chelonia mydas, at a temperate foraging habitat in the northern Gulf of Mexico: assessing short-term effects of cold stunning on growth.  Journal of Natural History 42:2793-2307.

Manire C.A, B.A. Stacy, M.J. Kinsel, H.T. Daniel, E.T. Anderson, and J.F. Wellehan Jr. 2008. Proliferative dermatitis in a loggerhead turtle, Caretta caretta, and a green turtle, Chelonia mydas, associated with novel papillomaviruses. Vet Microbiol. 130(3-4):227-37.

National Marine Fisheries Service and U.S. Fish and Wildlife Service. 2008. Recovery Plan for the Northwest Atlantic Populations of the Loggerhead Sea Turtle (Caretta caretta). National Marine Fisheries Service, Silver Spring, MD. Team: A.B. Bolten (team leader), L.B. Crowder, M.G. Dodd, S.L. MacPherson, J.A. Musick, B.A. Schroeder, and B.E. Witherington. 323 pages.

Pfaller, J.B., C.J. Limpus, and K.A. Bjorndal. 2008. Nest-site selection in individual loggerhead turtles and consequences for doomed-egg relocation. Conservation Biology 23:72–80. Pfaller_et_al_ConsBiol_2008.pdf

Pfaller, J.B., M.G. Frick, K.J. Reich, K.L. Williams, and K.A. Bjorndal. 2008. Carapace epibionts of loggerhead turtles (Caretta caretta) nesting at Canaveral National Seashore, Florida. Journal of Natural History 42:1095–1102. Pfaller_et_al_JNatHist_2008.pdf

Reich, K.J., K.A. Bjorndal, and C. Martínez del Rio. 2008. Effects of growth and tissue type on the kinetics of 13C and 15N incorporation in a rapidly growing ectotherm. Oecologia 155:651-663. Reich_et_al_Oecologia_2008.pdf

Stacy, B.A, M. Santoro, J.A. Morales, L.M. Huzella, V.F. Kalasinsky, A. Foley, N. Mettee, and E.R. Jacobson. 2008. Renal oxalosis in free-ranging green turtles Chelonia mydas. Dis Aquat Organ 80(1):45-9.

Stacy B.A, J.F. Wellehan, A.M. Foley, S.S. Coberley, L.H. Herbst, C.A. Manire, M.M Garner, M.D. Brookins, A.L. Childress, and E.R. Jacobson. 2008. Two herpesviruses associated with disease in wild Atlantic loggerhead sea turtles (Caretta caretta). Vet Microbiol.126(1-3):63-73.


Bolker, B.M., T. Okuyama, K.A. Bjorndal, and A.B. Bolten. 2007. Incorporating multiple mixed stocks in mixed stock analysis: “many-to-many” analyses. Molecular Ecology 16:685-695. Bolker_et_al_MolecEcol_2007.pdf

Bowen, B.W., W.S. Grant, Z. Hillis-Starr, D.J. Shaver, K.A. Bjorndal, A.B. Bolten, and A.L. Bass. 2007. The advocate and the scientist: debating the commercial exploitation of endangered hawksbill turtles. Molecular Ecology 16:3514–3515. Bowen_et_al_MolecEcol_2007_response.pdf

Bowen, B.W., W.S. Grant, Z. Hillis-Starr, D.J. Shaver, K.A. Bjorndal, A.B. Bolten, and A.L. Bass. 2007. Mixed-stock analysis reveals the migrations of juvenile hawksbill turtles (Eretmochelys imbricata) in the Caribbean Sea. Molecular Ecology 16:49–60. Bowen_et_al_2007.pdf

Moran, K.L. and K.A. Bjorndal. 2007. Simulated green turtle grazing affects nutrient composition of the seagrass Thalassia testudinum. Marine Biology 150:1083-1092. Moran_&_Bjorndal_MarBiol_2007.pdf

Reich, K.J., K.A. Bjorndal, and A.B. Bolten. 2007. The ‘lost years’ of green turtles: using stable isotopes to study cryptic lifestages. Biology Letters 3:712–714. Reich_et_al_BiolLett_2007.pdf

Santos, M.A., A.B. Bolten, H.R. Martins, B. Riewald, and K.A. Bjorndal. 2007. Air-breathing visitors to seamounts: Section B: Sea turtles. Pages 239-244 in Pitcher, T.J., T. Morato, P.J.B. Hart, M.R. Clark, N. Haggan, and R.S. Santos, editors, Seamounts: ecology, fisheries and conservation. Blackwell Publishing, London.

Seminoff, J.A., K.A. Bjorndal, and A.B. Bolten. 2007. Stable carbon and nitrogen isotope discrimination and turnover in pond sliders Trachemys scripta: insights for trophic study of freshwater turtles. Copeia 2007:534–542. Seminoff_etal_2007_Ts_isotopes_Copeia.pdf

Strik (Stacy), N.I., A.R. Alleman, and K. Harr.  2007.  Comparative reptile hematology – circulating inflammatory cells.  In E.R. Jacobson (ed.) Infectious diseases and pathology of reptiles: A color atlas and text. CRC Press.


Bjorndal, K.A., A.B. Bolten, L. Moreira, C. Bellini, and M.Â. Marcovaldi. 2006. Population structure and diversity of Brazilian green turtle rookeries based on mitochondrial DNA sequences. Chelonian Conservation and Biology 5:262-268. Bjorndal_et_al_CCB_2006.pdf

Bolten, A.B. and K.A. Bjorndal. 2006. Movements of an olive ridley sea turtle (Lepidochelys olivacea) in The Bahamas. The Bahamas Naturalist and Journal of Science 1:59-60.

Bouchard, S.S. and K.A. Bjorndal. 2006. Ontogenetic diet shifts and digestive constraints in the omnivorous freshwater turtle Trachemys scripta. Physiological and Biochemical Zoology 79:150-158. Bouchard&Bjorndal_PBZ2006.pdf

Bouchard, S.S. and K.A. Bjorndal. 2006. An investigation of nonadditive interactions between animal and plant diet items in an omnivorous freshwater turtle Trachemys scripta. Comparative Biochemistry and Physiology 144:77-85. Bouchard & Bjorndal CBP 2006.pdf

Carthy, R.R. 2006. An owner’s manual for the Chelonioidea. Conservation Biology. 20(1)259-260.

Harrison, A.L. and K.A. Bjorndal. 2006. Connectivity and wide-ranging species in the ocean. Pages 213-232 in K.R. Crooks and M.A. Sanjayan (editors), Connectivity Conservation. Cambridge University Press, Cambridge.

Jacobson,E.R., Homer,B., Greiner,E., Szabo,N.J., Chrisman,C., Foley,A., Landsberg, J.H., Flewelling,L., Ewing,R., Moretti,R., Schaf,S., Rose,C., Mader,D., Harman,G., Manire,C., Mettee,N., Mizisin,A.P., Shelton, G.D. 2006. A neurological disease of loggerhead sea turtles (Caretta caretta) in coastal waters of Florida, USA. Diseases of Aquatic Organisms 70:139-154. Jacobson_et_al_DAO_2006.pdf

Moncada, F., F.A. Abreu-Grobois, A. Muhlia-Melo, C. Bell, S. Troëng, K.A. Bjorndal, A.B. Bolten, A.B. Meylan, J. Zurita, G. Espinosa, G. Nodarse, R. Márquez-Millán, A. Foley, and L. Ehrhart. 2006. Movement patterns of green turtles (Chelonia mydas) in Cuba and adjacentCaribbean waters inferred from flipper tag recaptures. Journal of Herpetology 40:22–34. Moncada_et_al_JHerp2006.pdf

Pfaller, J.B., K.A. Bjorndal, K.J. Reich, K.L. Williams, and M.G. Frick. 2006. Distribution patterns of epibionts on the carapace of loggerhead turtles, Caretta caretta. JMBA Biodiversity Records, published online, 5381:1-4. Pfaller_et_al_Biodiversity_Notes_2006.pdf

Tiwari, M., K.A. Bjorndal, A.B. Bolten, and B.M. Bolker. 2006. Evaluation of density-dependent processes and green turtle, Chelonia mydas, hatchling production at Tortuguero, Costa Rica. Marine Ecology Progress Series 326:283-293. Tiwari_et_al_2006_MEPS.pdf


Bjorndal, K.A. 2005. Ciclo de vida longo, estudos a longo prazo [Long-lived sea turtles require long-term studies]. Revista do Tamar 10:4.

Bjorndal, K.A., A.B. Bolten, and M.Y. Chaloupka. 2005. Evaluating trends in abundance of immature green turtles, Chelonia mydas, in the Greater Caribbean. Ecological Applications 15:304-314. Bjorndal_et_al_EcolApplic_2005.pdf

Bjorndal, K.A., A.B. Bolten, and S. Troëng. 2005. Population structure and genetic diversity in green turtles nesting at Tortuguero, Costa Rica, based on mitochondrial DNA control region sequences. Marine Biology 147:1449-1457. Bjorndal_et_al_MarBiol2005.pdf

Bouchard, S.S. and K.A. Bjorndal. 2005. Microbial fermentation in juvenile and adult pond slider turtles, Trachemys scripta. Journal of Herpetology 39:321-324. Bouchard&Bjorndal_JHerpetol2005.pdf

Jacobson, E., Gronwall, R., Maxwell, L., Merrit, K., Harman, G. 2005.Plasma concentrations of enrofloxacin after single dose oral administration in loggerhead sea turtles, Caretta caretta . J Zoo Wildl Med. 36: 628-634.

Moran, K.L. and K.A. Bjorndal. 2005. Simulated green turtle grazing affects structure and productivity of seagrass pastures. Marine Ecology Progress Series 305:235-247. Moran&Bjorndal_MEPS2005.pdf

Okuyama, T., and B. M. Bolker. 2005. Combining genetic and ecological data to estimate sea turtle origins. Ecological Applications 15(1): 315-325. Okuyama&Bolker_EcolApplic2005.pdf

Tiwari, M., K.A. Bjorndal, A.B. Bolten, and B.M. Bolker. 2005. Intraspecific application of the mid-domain effect model: spatial and temporal nest distributions of green turtles, Chelonia mydas, at Tortuguero, Costa Rica. Ecology Letters 8:918–924. Tiwari_et_al_EcolLett_2005.pdf


Bolten, A.B. and M. Chaloupka. 2004. West Atlantic Green Turtle Population Modeling Workshop. Marine Turtle Newsletter 103:17. Bolten&Chaloupka_MTN2004.pdf

Bolten, A.B., H.R. Martins, E. Isidro, M. Santos, R. Ferreira, E. Bettencourt, A. Giga, A. Cruz, and K.A. Bjorndal. 2004. Experiment to evaluate gear modification on rates of sea turtle bycatch in the swordfish longline fishery in the Azores – Phase 1 and Phase 2. Pages 139-153 in K.J. Long and B.A. Schroeder (editors), Proceedings of the International Technical Expert Workshop on Marine Turtle Bycatch in Longline Fisheries,Seattle, Washington, 11-13 February 2003. NOAA Technical Memorandum NMFS-OPR-26.

Bowen, B.W., A.L. Bass, S.M. Chow, M. Bostrom, K.A. Bjorndal, A.B.. Bolten, T. Okuyama, B.M. Bolker, S. Epperly, E. Lacasella, D. Shaver, M. Dodd, S.R. Hopkins-Murphy, J.A. Musick, Mark Swingle, K. Rankin-Baransky, W. Teas, W.N. Witzell and P.H. Dutton. 2004. Natal homing in juvenile loggerhead turtles (Caretta caretta). Molecular Ecology 13:3797-3808. Bowen_et_al_MolecEcol2004.pdf

Frick, M.G., K.L. Williams, A.B. Bolten, K.A. Bjorndal, and H.R. Martins. 2004. Diet and fecundity of Columbus crabs Planes minutusassociated with oceanic-stage loggerhead sea turtles Caretta caretta and inanimate flotsam. Journal of Crustacean Biology 24:350-355. Frick_et_al_JCrustBiol2004.pdf

Herbst, L. H., A. Ene, M. Su, R. DeSalle, and J. Lenz. 2004. Tumor outbreaks in marine turtles are not due to recent herpesvirus mutations. Current Biology14, no. 17: R697-R699.


Baptistotte, C., J.C.A. Thomé, and K.A. Bjorndal. 2003. Reproductive biology and conservation status of the loggerhead sea turtle (Caretta caretta) in Espírito Santo, Brazil. Chelonian Conservation and Biology 4:523-529. Baptistotte_et_al_CCB_2003.pdf

Bjorndal, K.A. 2003. Foreword to the book Sea Turtles of the World by Douglas Perrine, Voyageur Press, Stillwater, MN.

Bjorndal, K.A. 2003. Roles of loggerhead sea turtles in marine ecosystems. Pages 235-254 in A.B. Bolten and B.E. Witherington (editors), Loggerhead sea turtles. Smithsonian Institution Press, Washington, D.C.

Bjorndal, K.A. and A.B. Bolten. 2003. From ghosts to key species: restoring sea turtle populations to fulfill their ecological roles. Invited paper for the 100th Issue of the Marine Turtle Newsletter 100:16-21. Bjorndal&Bolten_MTN100.pdf

Bjorndal, K.A., A.B. Bolten, A. Arenas, J. Zurita, A. D’Amiano, C. Calderón, J. Parsons, and J.A. Seminoff. 2003. Green turtle with living tag captured in the southern Bahamas. Marine Turtle Newsletter 101:26. Bjorndal_et_al_MTN2003.pdf

Bjorndal, K.A., A.B. Bolten, and M.Y. Chaloupka. 2003. Survival probability estimates for immature green turtles, Chelonia mydas, in theBahamas. Marine Ecology Progress Series 252:273-281. Bjorndal_et_al_MEPS2003.pdf

Bjorndal, K.A., A.B. Bolten, T. Dellinger, C. Delgado, and H.R. Martins. 2003. Compensatory growth in oceanic loggerhead sea turtles: response to a stochastic environment. Ecology 84:1237-1249. Bjorndal_et_al_Ecology2003.pdf

Bjorndal, K.A., A.B. Bolten, and H.R. Martins. 2003. Estimates of survival probabilities for oceanic-stage loggerhead sea turtles (Caretta caretta) in the North Atlantic. Fishery Bulletin 101:732-736. Bjorndal_et_al_FishBull2003.pdf

Bjorndal, K.A. and J.B.C. Jackson. 2003. Roles of sea turtles in marine ecosystems: reconstructing the past. Pages 259-273 in P.L. Lutz, J. Musick and J. Wyneken (editors), Biology of Sea Turtles, volume II. CRC Press, Boca Raton, FL.

Bolker, B., T. Okuyama, K. Bjorndal, and A. Bolten. 2003. Sea turtle stock estimation using genetic markers: accounting for sampling error of rare genotypes. Ecological Applications 13:763-775. Bolker_et_al_EcolApplic_2003.pdf

Bolten, A.B. 2003. Variation in sea turtle life history patterns: neritic vs. oceanic developmental stages. Pages 243-257 in P.L. Lutz, J. Musick and J. Wyneken (editors), The Biology of Sea Turtles, volume II. CRC Press, Boca Raton, FL. Bolten_chapter9(CRC Press).pdf

Bolten, A.B. 2003. Active swimmers – passive drifters: the oceanic juvenile stage of loggerheads in the Atlantic system. Pages 63-78 in A.B. Bolten and B.E. Witherington (editors), Loggerhead Sea Turtles. Smithsonian Institution Press, Washington, D.C. Bolten_Chapter4(Smithsonian Press).pdf

Bolten, A.B. 2003. The loggerhead sea turtle: a most excellent fishe. Pages 1-3 in A.B. Bolten and B.E. Witherington (editors). Loggerhead SeaTurtles. Smithsonian Institution Press, Washington, D.C.

Bolten, A.B. and K.A. Bjorndal. 2003. Green turtles in the Caribbean: a shared resource. Pages 225-226 in G.C. Ray and J. McCormick-Ray. Coastal-Marine Conservation: Science and Policy. Blackwell Scientific, London. Case Study within a chapter on the Bahamas.

Bolten, A.B. and B.E. Witherington (editors). 2003. Loggerhead sea turtles. Smithsonian Institution Press, Washington, D.C. 319 pages.

Carthy, R.R., A.M. Foley, and Y. Matsuzawa. 2003. The incubation environment of nests of the loggerhead turtle (Caretta caretta) and effects on hatching success and the characteristics of hatchlings. In “The Biology of the Loggerhead Sea Turtle”. A.B. Bolten and B. Witherington, eds. Smithsonian Institution Press, Washington D.C.

Frick, M.G., A. Ross, K.L. Williams, A.B. Bolten, K.A. Bjorndal, and H.R. Martins. 2003. Epibiotic associates of oceanic-stage loggerhead turtles from the southeastern North Atlantic. Marine Turtle Newsletter 101:18-20. Frick_et_al_MTN2003.pdf

Herbst, L. H., and E. R. Jacobson. 2003. Practical approaches for studying sea turtle health and disease. Editors P. L. Lutz, J. A. Musick, and J. Wyneken. The Biology of Sea Turtles. Volume II. CRC Marine Biology Series, CRC Press, Inc.: Boca Raton, London, New York, Washington D.C. 455 Pp. p.385-410.

Herbst, L.H., Siconolfi-Baez, L., P.A. Klein, M.J. Kerben, I.M. Schumacher. 2003. Induction of vitellogenin by Estradiol-17beta and development of enzyme linked immunosorbant assays to quantify plasma vitellogenin levels in green turtles (Chelonia mydas). Comp Biochem Physiol B Biochem Mol Biol. 135(3): 551-63.

Jacobson, E., Harman, G., Maxwell, L., and Laille, E. 2003. Plasma concentrations of Praziquantel in loggerhead sea turtles, Caretta caretta, following oral administration of single and multiple doses. Amer J Vet Res. 64: 304-308.

Pandolfi, J.M., R.H. Bradbury, E. Sala, T.P. Hughes, K.A. Bjorndal, R.G. Cooke, D. McArdle, L. McClenachan, M.J.H. Newman, G. Paredes, R.R. Warner, and J.B.C. Jackson. 2003. Global trajectories of the long-term decline of coral reef ecosystems. Science 301:955-958. Pandolfi_et_al_Science2003.pdf

Schmid, J.R., A.B. Bolten, K.A. Bjorndal, W.J. Lindberg, H.F. Percival, and P.D. Zwick. 2003. Home range and habitat use by Kemp’s ridley turtles in west-central Florida. Journal of Wildlife Management 67:196-206.

Tiwari, M., R.R. Carthy, A. Silveira, and K.A. Bjorndal. 2003. Caretta caretta (loggerhead sea turtle). Nest architecture. Herpetological Review 34:138-139.


Coberley, S.S., Condit, R.C., Herbst, L.H., and P. A. Klein. 2002. Identification and Expression of Immunogenic Proteins of a Disease-associated Marine Turtle Herpesvirus. J. Virol. 76: 10553-10558.

León, Y.M. and K.A. Bjorndal. 2002. Selective feeding in the hawksbill turtle, an important predator in coral reef ecosystems. Marine Ecology Progress Series 245:249-258. Leon&Bjorndal_MEPS2002.pdf

Matsuzawa, Y., K. Sato, W. Sakamoto, and K.A. Bjorndal. 2002. Seasonal fluctuations in sand temperature: effects on the incubation period and mortality of loggerhead sea turtle (Caretta caretta) pre-emergent hatchlings in Minabe, Japan. Marine Biology 140:639-646. Matsuzawa_et_al_MarBiol2002.pdf

Maglothin, M.L., M.M. Lamont, and R.R. Carthy. 2002. Nest Chamber anomalies in loggerhead sea turtle (Caretta caretta) nests from Northwest Florida, USA. Herpetological Review. 33(1):48-50.

Parris, L., M.M. Lamont, and R.R. Carthy. 2002. Observations of red imported fire ant predation on loggerhead turtle hatchlings. FloridaEntomologist. 85(3):514-517.

Schmid, J.R., A.B. Bolten, K.A. Bjorndal, and W.J. Lindberg. 2002. Activity patterns of Kemp’s ridley turtles, Lepidochelys kempii, in the coastal waters of the Cedar Keys, Florida. Marine Biology 140:215-228. Schmid_et_al_MarBiol2002.pdf

Solow, A.R., K.A. Bjorndal, and A.B. Bolten. 2002. Annual variation in nesting numbers of marine turtles: the effect of sea surface temperature on re-migration intervals. Ecology Letters 5:742-746. Solow_et_al_EcolLet2002.pdf


Bjorndal, K.A., A.B. Bolten, B. Koike, B.A. Schroeder, D.J. Shaver, W.G. Teas, and W.N. Witzell. 2001. Somatic growth function for immature loggerhead sea turtles in southeastern U.S. waters. Fishery Bulletin 99:240-246. Bjorndal_et_al_FishBull2001.pdf

Coberley, S. S., Herbst, L. H., Brown, D. R., Ehrhart, L. M., Bagley, D. A., Schaf, S., Moretti, R., Jacobson, E. R., Klein, P. A. 2001. Detection of Antibodies to a Disease-Associated Herpesvirus of the Green Turtle, Chelonia mydas. Journal of Clinical Microbiology 39:3572-3577.

Coberley, S.S., Herbst, L.H., Ehrhart, L.M., Bagley, D.A., Hirama, S., Jacobson, E.R., Klein, P.A. 2001. Survey of Florida Green Turtles for Exposure to a Disease-Associated Herpesvirus. Diseases of Aquatic Organsims 47:159-167.

Ferreira, R.L., H.R. Martins, A.A. da Silva, and A.B. Bolten. 2001. Impact of swordfish fisheries on sea turtles in the Azores. Arquipélago 18A:75-79.

Herbst, L. H., Chakrabarti, R., Klein, P. A., and Achary, M. 2001. Differential gene expression associated with tumorigenicity of cultured green turtle fibropapilloma-derived fibroblasts. Cancer Genetics and Cytogenetics129, no. 1: 35-39.

Jackson, J. B. C., M. X. Kirby, W. H. Berger, K. A. Bjorndal, L. W. Botsford, B. J. Bourque, R. H. Bradbury, R. Cooke, J. Erlandson, J. A. Estes, T. P. Hughes, S. Kidwell, C. B. Lange, H. S. Lenihan, J. M. Pandolfi, C. H. Peterson, R. S. Steneck, M. J. Tegner, and R. R. Warner. 2001. Historical overfishing and the recent collapse of coastal ecosystems. Science 293:629-638. Jackson_et_al_Science2001.pdf

Martins, H.R., A.B. Bolten, and R.L. Ferreira. 2001. Impact of longline fisheries on sea turtles in the archipelago of the Azores. Pages 76-77 in S. Ciccione, D. Ross, and J. Le Gall, editors. Advance in knowledge and conservation of sea turtles in south-west Indian Ocean. Etudes et Colloques du Editions du Centre d’Etude et de Découverte des Tortues Marine de La Réunion (CEDTM).

Oros, J., Torrent, A., De Los Monteros, E., Calabuig, P., Deniz, S., Tucker, S., Jacobson, E.R. 2001. Multicentric lymphoblastic Lymphoma In A Loggerhead Sea Turtle (Caretta caretta). Vet. Path. 38:464 467.


Bjorndal, K.A. and A.B. Bolten (editors). 2000. Proceedings of a workshop on assessing abundance and trends for in-water sea turtle populations. NOAA Technical Memorandum NMFS-SEFSC-445. 83 pages.

Bjorndal, K.A. and A.B. Bolten. 2000. Green turtles at Conception Island Creek, Bahamas. Pages 75-76 in K.A. Bjorndal and A.B. Bolten (editors), Proceedings of a workshop on assessing abundance and trends for in-water sea turtle populations. NOAA Technical Memorandum NMFS-SEFSC-445. Bjorndal&Bolten_Pop_Assess_Workshop2000.pdf

Bjorndal, K.A., A.B. Bolten, and M.Y. Chaloupka. 2000. Green turtle somatic growth model: evidence for density dependence. Ecological Applications 10:269-282. Bjorndal_et_al_EcolApplic2000.pdf

Bjorndal, K.A., A.B. Bolten, and H.R. Martins. 2000. Somatic growth model of juvenile loggerhead sea turtles Caretta caretta: duration of pelagic stage. Marine Ecology Progress Series 202:265-272. Bjorndal_et_al_MEPS2000.pdf

Bolten, A.B., H.R. Martins, and K.A. Bjorndal (editors). 2000. Proceedings of a workshop to design an experiment to determine the effects of longline gear modification on sea turtle bycatch rates [Workshop para a elaboração de uma experiência que possa diminuir as capturas acidentais de tartarugas marinhas nos Açores], 2-4 September 1998, Horta, Azores, Portugal. NOAA Technical Memorandum NMFS-OPR-19. 50 pages. Bilingual

Bouchard, S.S. and K.A. Bjorndal. 2000. Sea turtles as biological transporters of nutrients and energy from marine to terrestrial ecosystems. Ecology 81:2305-2313. Bouchard&Bjorndal_Ecology2000.pdf

Curry, S., Brown, D. R., Gaskin, J. M., Jacobson, E. R., Ehrhart, L. M., Blahak, S., Herbst, L. H., and P. A. Klein. 2000. Persistent infectivity of a green turtle disease-associated herpesvirus after exposure to seawater. J. Wildlife Diseases, 36, 2000, 792-797.

Eliazar, P.J., K.A. Bjorndal, and A.B. Bolten. 2000. Early report of fibropapilloma from St. Croix, USVI. Marine Turtle Newsletter 89:16. Eliazar_et_al_MTN2000.pdf

Tiwari, M. and K.A. Bjorndal. 2000. Variation in morphology and reproduction in loggerheads, Caretta caretta, nesting in the United States,Brazil, and Greece. Herpetologica 56:343-356.

Wood, D.W. and K.A. Bjorndal. 2000. Relation of temperature, moisture, salinity, and slope to nest site selection in loggerhead sea turtles. Copeia 2000:119-128. Wood&Bjorndal_Copeia2000.pdf


Bjorndal, K.A. 1999. Priorities for research in foraging habitats. Pages 12-14 in K.L. Eckert, K.A. Bjorndal, F.A. Abreu Grobois and M. Donnelly (editors). Management and Research Techniques for the Conservation of Sea Turtles. IUCN, Gland, Switzerland. Bjorndal_Tech_Man_1999.pdf

Bjorndal, K.A. 1999. Conservation of hawksbill sea turtles: perceptions and realities. Chelonian Conservation and Biology 3:174-176. Bjorndal_CCB_1999.pdf

Bjorndal, K.A., J.A. Wetherall, A.B. Bolten, and J.A. Mortimer. 1999. Twenty-six years of green turtle nesting at Tortuguero, Costa Rica: an encouraging trend. Conservation Biology 13:126-134. Bjorndal_et_al_ConsBio_1999.pdf

Bolten, A.B. 1999. Techniques for measuring sea turtles. Pages 110-114 in K.L. Eckert, K.A. Bjorndal, F.A. Abreu Grobois and M. Donnelly (editors). Management and Research Techniques for the Conservation of Sea Turtles. IUCN, Gland, Switzerland. Bolten_Tech_Man_1999.pdf

Eckert, K.L., K.A. Bjorndal, F.A. Abreu Grobois and M. Donnelly (editors). 1999. Management and Research Techniques for the Conservation of Sea Turtles. IUCN, Gland, Switzerland. 235 pages. Spanish translation published in 2001. Técnicas de Investigación y Manejo para la Conservación de las Tortugas Marinas. 270 pages.

Herbst, L. H., Jacobson, E. R., Klein, P. A., Balazs, G. H., Moretti, R. Brown, T., and Sundberg, J. P. 1999. Comparative pathology and pathogenesis of experimentally induced and spontaneous fibropapillomas of green turtles (Chelonia mydas) Veterinary Pathology 36(6): 551-564.

Jacobson, E.R. 1999. Necropsy and tissue sampling techniques for marine turtles. In: Eckert, K., Bjorndal, K., and Abreu, A. (Eds). Management and Research Techniques for the Conservation of Sea Turtles. I.U.C.N. Pp. 214-217.

Jacobson, E.R., Behler, J.L., Jarchow, J.L. 1999. Health assessment of chelonians and release into the wild. In: Zoo and Wild Animal Medicine, Current Therapy 4. Fowler ME and Miller RE (eds), WB Saunders Co, Philadelphia PA, Pp. 232-242.

Lackovich, J.K., Brown, D.R., Homer, B.L., Garber, R.L., Mader, D.R., Moretti, R.H., Patterson, A.D., Herbst, L.H.. Oros, J., Jacobson, E.R., Curry, S.S., Klein, P.A. 1999, Association of herpesvirus with fibropapillomatosis of the green turtle, Chelonia mydas, and the loggerhead turtle, Caretta caretta, in Florida. Dis. Aq. Org. 37: 89 97.

Lackovich, J., Brown, D.R., and Klein, P.A. 1999.Further evidence for absence of papillomaviruses from sea turtle fibropapillomas. Veterinary Record 145: 616-617.

McCauley, S.J. and K.A. Bjorndal. 1999. Conservation implications of dietary dilution from debris ingestion: sublethal effects in post-hatchling loggerhead sea turtles. Conservation Biology 13:925-929. McCauley&Bjorndal_ConsBiol1999.pdf

McCauley, S.J. and K.A. Bjorndal. 1999. Response to dietary dilution in an omnivorous freshwater turtle: implications for ontogenetic dietary shifts. Physiological and Biochemical Zoology 72:101-108. McCauley&Bjorndal_PBZ1999.pdf

Moran, K.L., K.A. Bjorndal, and A.B. Bolten. 1999. Effects of the thermal environment on the temporal pattern of emergence of hatchling loggerhead turtles (Caretta caretta). Marine Ecology Progress Series 189:251-261.


Bjorndal, K.A. and A.B. Bolten. 1998. Hawksbill tagged in The Bahamas recaptured in Cuba. Marine Turtle Newsletter 79:18-19. Bjorndal&Bolten_MTN1998.pdf

Bjorndal, K.A., A.B. Bolten, R.A. Bennett, E.R. Jacobson, T.J. Wronski, J.J. Valeski, and P.J. Eliazar. 1998. Age and growth in sea turtles: limitations of skeletochronology for demographic studies. Copeia 1998:23-30.

Bolten, A.B., K.A. Bjorndal, H.R. Martins, T. Dellinger, M.J. Biscoito, S.E. Encalada, and B.W. Bowen. 1998. Transatlantic developmental migrations of loggerhead sea turtles demonstrated by mtDNA sequence analysis. Ecological Applications 8:1-7. Bolten_et_al_EcolApplic_1998.pdf

Bouchard, S., K. Moran, M. Tiwari, D. Wood, A. Bolten, P. Eliazar, and K. Bjorndal. 1998. Effects of exposed pilings on sea turtle nesting activity at Melbourne Beach, Florida. Journal of Coastal Research 14:1343-1347.

Encalada, S.E., K.A. Bjorndal, A.B. Bolten, J.C. Zurita, B. Schroeder, E. Possardt, C.J. Sears, and B.W. Bowen. 1998. Population structure of loggerhead turtle (Caretta caretta) nesting colonies in the Atlantic and Mediterranean as inferred from mitochondrial DNA control region sequences. Marine Biology 130:567-575. Encalada_et_al_MarBiol1998.pdf

Herbst. L. H., Greiner, E. C., Ehrhart, L. M., Bagley, D. A., and P. A. Klein. 1998.Serologic association between spirorchidiasis, herpesvirus infection, and fibropapillomatosis in green turtles (Chelonia mydas) from Florida. J. Wildlife Diseases 34: 496-507.

Herbst, L.H., Sundberg, J.P., Shultz, L.D, Gray, B.A., and P. A. Klein. 1998.Tumorigenicity of green turtle fibropapilloma-derived fibroblast lines in immunodeficient mice. Laboratory Animal Science 48(2): 162-167.

Jacobson, E.R. 1998. Collecting and processing blood from sea turtles for hematologic and plasma biochemical determinations. In: report of the Sea Turtle Health Assessment Workshop, 2-3 February 1998. Pat I: Background and Information Needs, Fair PA and Hansen LJ (eds). NOAA Technical Memorandum NOS-NCCOS-CCEHBR-0003, Pp.24-31.

Lahanas, P.N, K.A. Bjorndal, A.B. Bolten, S.E. Encalada, M.M. Miyamoto, R.A.Valverde, and B.W. Bowen. 1998. Genetic composition of a green turtle (Chelonia mydas) feeding ground population: evidence for multiple origins. Marine Biology 130:345-352. Lahanas_et_al_MarBiol1998.pdf


Bjorndal, K.A. 1997. Foraging ecology and nutrition of sea turtles. Pages 199-231 in P.L. Lutz and J.A. Musick (eds.), The biology of sea turtles. CRC Press, Boca Raton, Florida.

Bjorndal, K.A. 1997. Fermentation in reptiles and amphibians. Pages 199-230 in R.I. Mackie and B.A. White (eds.), Gastrointestinal microbiology, Vol. 1 Gastrointestinal ecosystems and fermentations. Chapman and Hall, New York.

Bjorndal, K.A., A.B. Bolten, C.J. Lagueux, and D.R. Jackson. 1997. Dietary overlap in three sympatric congeneric freshwater turtles (Pseudemys) in Florida. Chelonian Conservation and Biology 2:430-433. Bjorndal_et_al_CCB_1997.pdf


Bass, A.L., D.A. Good, K.A. Bjorndal, J.I. Richardson, Z.-M. Hillis, J.A. Horrocks, and B.W. Bowen. 1996. Testing models of female reproductive migratory behaviour and population structure in the Caribbean hawksbill turtle, Eretmochelys imbricata, with mtDNA sequences. Molecular Ecology 5:321-328.

Bjorndal, K.A., A.B. Bolten, C.J. Lagueux, and A. Chaves. 1996. Probability of tag loss in green turtles nesting at Tortuguero, Costa Rica. Journal of Herpetology 30:566-571. Bjorndal_et_al_JHerpetol1996.pdf

Bolten, A.B., J.A. Wetherall, G.H. Balazs and S.G. Pooley (compilers). 1996. Status of marine turtles in the Pacific Ocean relevant to incidental take in the Hawaii-based pelagic longline fishery. NOAA Technical Memorandum NOAA-TM-NMFS-SWFSC-230. 167 pages.

Bowen, B.W., A.L. Bass, A. Garcia-Rodriguez, C.E. Diez, R. van Dam, A.B. Bolten, K.A. Bjorndal, M.M. Miyamoto, and R.J. Ferl. 1996. Origin of hawksbill turtles in a Caribbean feeding area as indicated by genetic markers. Ecological Applications 6:566-572. Bowen_et_al_EcolApplic1996.pdf

Dunn, F.P., K.A. Bjorndal, J.M. Coleman, W.E. Evans, R. Kasprzak, J.E. Kiesler, P. O’Connor, A. Powell, A.G. Pulsipher, D.J. Sullivan, J.P. VanDevender and P.K Vélez. 1996. An assessment of techniques for removing offshore structures. National Academy Press, Washington,D.C. 76 pages.

Encalada, S.E., P.N. Lahanas, K.A. Bjorndal, A.B. Bolten, M.M. Miyamoto, and B.W. Bowen. 1996. Phylogeography and population structure of the Atlantic and Mediterranean green turtle Chelonia mydas: a mitochondrial DNA control region sequence assessment. Molecular Ecology 5:473-483.

Gregory, L.F., T.S. Gross, A.B. Bolten, K.A. Bjorndal, and L.J. Guillette, Jr. 1996. Plasma corticosterone concentrations associated with acute captivity stress in wild loggerhead sea turtles (Caretta caretta). General and Comparative Endocrinology 104:312-320. Gregory_et_al_GenCompEndocrin1996.pdf

Herbst, L.H. and Klein, P.A. 1996. Analysis of tumorigenicity and differential gene expression in fibroblast lines derived from normal skin and fibropapillomas of the green turtle (Chelonia mydas) National Marine Fisheries Service, NOAA, Honolulu, Hawaii, Administrative report H-96-04C, June, 1996.

Herbst, L.H., Moretti, R., Brown, T., and P. A. Klein. 1996. Sensitivity of the transmissible green turtle fibropapillomatosis agent to chloroform and ultracentrifugation conditions. Diseases of Aquatic Organisms 25(3): 225-228.

Johnson, S.A., K.A. Bjorndal, and A.B. Bolten. 1996. Loggerhead turtle (Caretta caretta) nesting behavior and hatchling production in Florida: Effects of organized turtle watches. Conservation Biology 10:570-577. Johnson_et_al_ConsBio1996.pdf

Johnson, S.A., K.A. Bjorndal, and A.B. Bolten. 1996. A survey of organized turtle watch participants on sea turtle nesting beaches in Florida. Chelonian Conservation and Biology 2:60-65.

Schumacher, J., Papendick, R., Herbst, L., and Jacobson, E.R. 1996. Volvulus of the Proximal Colon in a Hawksbill Turtle (Eretmochelys imbricata). J. Zoo Wildlf. Med. 27: 386-391.


Bjorndal, K.A. (editor). 1995. Biology and conservation of sea turtles, revised edition. Smithsonian Institution Press, Washington, D.C. 615 pages.

Bjorndal, K.A. and A.B. Bolten. 1995. Comparison of length-frequency analyses for estimation of growth parameters for a population of green turtles. Herpetologica 51:160-167. Bjorndal_&_Bolten_Herpetologica_1995.pdf

Bjorndal, K.A., A.B. Bolten, A.L. Coan Jr., and P. Kleiber. 1995. Estimation of green turtle (Chelonia mydas) growth rates from length-frequency analysis. Copeia 1995:71-77. Bjorndal_et_al_Copeia_1995.pdf

Bjorndal, K.A. and G.R. Zug. 1995. Growth and age of sea turtles. Pages 599-600 in K.A. Bjorndal (ed.), Biology and conservation of sea turtles, revised edition. Smithsonian Institution Press, Washington, D.C.

Bolten, A.B. and G.H. Balazs. 1995. Biology of the early pelagic stage–the “lost year.” Pages 575-581 in K.A. Bjorndal (editor), Biology and Conservation of Sea Turtles, revised edition. Smithsonian Institution Press, Washington, D.C.

Bolten, A.B., K.A. Bjorndal and H.R. Martins. 1995. Life history of the loggerhead sea turtle, Caretta caretta (Reptilia: Cheloniidae), in theAtlantic. In A. Domingo Abreu and M. Biscoito (eds.), Proceedings of the First Symposium Fauna and Flora of Atlantic Islands. Bol. Museu Municipal do Funchal, Madeira, Portugal, Sup. no. 4:115-122.

Crain, D.A., A.B. Bolten, and K.A. Bjorndal. 1995. Effects of beach nourishment on sea turtles: Review and research initiatives. Restoration Ecology 3:95-104.

Crain, D.A., A.B. Bolten, K.A. Bjorndal, L.J. Guillette Jr,. and T.S. Gross. 1995. Size-dependent, sex-dependent, and seasonal changes in insulin-like growth factor I in the loggerhead sea turtle (Caretta caretta). General and Comparative Endocrinology 98:219-226. Crain_et_al_GenCompEndocrin1995.pdf

Gross, T.S., D.A. Crain, K.A. Bjorndal, A.B. Bolten, and R.R. Carthy. 1995. Identification of sex in hatchling loggerhead turtles (Caretta caretta) by analysis of steroid concentrations in chorioallantoic fluid. General and Comparative Endocrinology 99:204-210. Gross_et_al_GenCompEndocrin1995.pdf

Herbst, L. H., and E. R. Jacobson. 1995. Diseases of marine turtles. Editor K. A. Bjorndal. Biology and Conservation of Sea Turtles (Revised Edition). Smithsonian Institution Press, Washington, D.C. and London. 615 Pp. p.593-96.

Herbst, L.H., Jacobson, E.R., Moretti, R., Brown, T., Sundberg, J.P. and Klein, P.A. 1995. Experimental Transmission of Green Turtle Fibropapillomatosis Using Cell-free Extracts. Diseases of Aquatic Organisms 22(1):1-12.

Herbst, L.H. and P.A. Klein. 1995. Monoclonal antibodies for the measurement of class-specific antibody responses in the Green Turtle, Chelonia Mydas. J. Veterinary Immunology and Immunopathology. 46: 317-335.

Herbst , L.H. and Klein, P.A. 1995. Green Turtle Fibropapillomatosis: Challenges to assessing the role of environmental cofactors. Environmental Health Perspectives. 103: 27-30.

Lagueux, C.J., K.A. Bjorndal, A.B. Bolten, and C.L. Campbell. 1995. Food habits of Pseudemys concinna suwanniensis in a Florida spring. Journal of Herpetology 29:122-126. Lagueux_et_al_JHerpetol1995.pdf


Allard, M.W., M.M. Miyamoto, K.A. Bjorndal, A.B. Bolten, and B.W. Bowen. 1994. Support for natal homing in green turtles from mitochondrial DNA sequences. Copeia 1994:34-41. Allard_et_al_Copeia_1994_high_res.pdf

Bjorndal, K.A., A.B. Bolten, J. Gordon, and J.A. Camiñas. 1994. Caretta caretta (loggerhead) growth and pelagic movement. Herpetological Review 25:23-24.

Bjorndal, K.A., A.B. Bolten, D.A. Johnson, and P.J. Eliazar (compilers). 1994. Proceedings of the Fourteenth Annual Symposium on Sea Turtle Biology and Conservation. NOAA Technical Memorandum NMFS-SEFSC-351. 323 pages.

Bjorndal, K.A., A.B. Bolten, and C.J. Lagueux. 1994. Ingestion of marine debris by juvenile sea turtles in coastal Florida habitats. Marine Pollution Bulletin 28:154-158. BjorndalEtAl_1994_IngestionOfMarineDebrisByJuvenileSeaTurtlesInCoastalFlorida.pdf

Bolten, A.B., K.A. Bjorndal, P.J. Eliazar, and L.F. Gregory. 1994. Seasonal abundance, size distribution, and blood biochemical values of loggerheads (Caretta caretta) in Port Canaveral Ship Channel, Florida. NOAA Technical Memorandum NMFS-SEFSC-353. 39 pages.

Bolten, A.B., K.A. Bjorndal and H.R. Martins. 1994. Life history model for the loggerhead sea turtle (Caretta caretta) population in the Atlantic: Potential impacts of a longline fishery. Pages 48-55 in G.H. Balazs and S.G. Pooley (eds.), Research plan to assess marine turtle hooking mortality: Results of an expert workshop held in Honolulu, Hawaii November 16-18, 1993. NOAA Technical Memorandum NMFS-SWFSC-201.

Brooks, D.E., Ginn, P.E., Jacobson, E.R., Miller, T.R. and Bramson, L. 1994. Ocular Fibropapillomas of Green Turtles (Chelonia mydas). Vet. Path. 31:335-339.

Herbst, L. H. 1994. Fibropapillomatosis of marine turtles. Annual Review of Fish Diseases4: 389-425.

Homer, B.L., Jacobson, E.R., Schumacher, J. and Scherba, G. 1994. Chlamydiosis in Green Sea Turtles. Vet. Pathol. 31:1-7.

Lahanas, P.N., M.M. Miyamoto, K.A. Bjorndal, and A.B. Bolten. 1994. Molecular evolution and population genetics of Greater Caribbean green turtles (Chelonia mydas) as inferred from mitochondrial DNA control region sequences. Genetica 94:57-67.


Bjorndal, K.A. and A.B. Bolten. 1993. Digestive efficiencies in herbivorous and omnivorous freshwater turtles on plant diets: do herbivores have a nutritional advantage? Physiological Zoology 66:384-395.

Bjorndal, K.A., A.B. Bolten, and C.J. Lagueux. 1993. Decline of the nesting population of hawksbill turtles at Tortuguero, Costa Rica. Conservation Biology 7:925-927. Bjorndal_et_alConsBio_1993.pdf

Bolten, A.B., H.R. Martins, K.A. Bjorndal, and J. Gordon. 1993. Size distribution of pelagic-stage loggerhead sea turtles (Caretta caretta) in the waters around the Azores and Madeira. Arquipélago 11A:49-54. Bolten_et_al_Arqui_1993.pdf

Klein, P.A., L. Herbst, E. Jacobson, A.B. Bolten, B. R. Collins, and E. Greiner. 1993. Development of immunodiagnostic tools for studying the etiology and epidemiology of green turtle fibropapillomatosis. National Marine Fisheries Service, NOAA, Honolulu, Hawaii, Administrative report H-93-13C, July, 1993.


Bjorndal, K.A. and A.B. Bolten. 1992. Spatial distribution of green turtle (Chelonia mydas) nests at Tortuguero, Costa Rica. Copeia 1992:45-53. Bjorndal&Bolten_Copeia1992.pdf

Bjorndal, K.A. and A.B. Bolten. 1992. Body size and digestive efficiency in a herbivorous freshwater turtle: advantages of small bite size. Physiological Zoology 65:1028-1039.

Bolten, A.B. and K.A. Bjorndal. 1992. Blood profiles for a wild population of green turtles (Chelonia mydas) in the southern Bahamas: size-specific and sex-specific relationships. Journal of Wildlife Diseases 28:407-413.

Bolten, A.B., K.A. Bjorndal, J.S. Grumbles, and D.W. Owens. 1992. Sex ratio and sex-specific growth rates in immature green turtles,Chelonia mydas, in the southern Bahamas. Copeia 1992:1098-1103. Bolten_et_al_Copeia_1992.pdf

Bolten, A.B., E.R. Jacobson, and K.A. Bjorndal. 1992. Effects of anticoagulant and autoanalyzer on blood biochemical values of loggerhead sea turtles (Caretta caretta). American Journal of Veterinary Research 53:2224-2227.

Bolten, A.B., H.R. Martins, K.A. Bjorndal, M. Cocco, and G. Gerosa. 1992. Caretta caretta (loggerhead) pelagic movement and growth. Herpetological Review 23:116.

Bolten, A.B., J.C. Santana, and K.A. Bjorndal. 1992. Transatlantic crossing by a loggerhead turtle. Marine Turtle Newsletter 59:7-8.


Bjorndal, K.A. 1991. Preservation of sea turtles and international cooperation. Pages 11-14 in I. Uchida (ed.), Proceedings of the International Symposium on Sea Turtles ’88. Himeji City Aquarium and Hiwasa Chelonian Museum, Japan.

Bjorndal, K.A. 1991. Diet mixing: non-additive interactions of diet items in an omnivorous freshwater turtle. Ecology 72:1234-1241. Bjorndal_Ecology_1991.pdf

Bjorndal, K.A., H. Suganuma, and A.B. Bolten. 1991. Digestive fermentation in green turtles, Chelonia mydas, feeding on algae. Bulletin of Marine Science 48:166-171. BjorndalEtAl_1991_DigestiveFermentationInGreenTurtles.pdf

Guillette, L.J., Jr., K.A. Bjorndal, A.B. Bolten, T.S. Gross, B.D. Palmer, B.E. Witherington, and J.M. Matter. 1991. Plasma estradiol-17B, progesterone, prostaglandin F, and prostaglandin E2 concentrations during natural oviposition in the loggerhead turtle (Caretta caretta). General and Comparative Endocrinology 82:121-130.

Jacobson, E.R., Buergelt, C., Williams, B. and Harris, R.K. 1991. Herpesvirus in Cutaneous Fibropapillomas of the Green Turtle Chelonia mydas. Dis. Aquatic Org. 12:1-6.

Manzella, S., K. Bjorndal, and C. Lagueux. 1991. Head-started Kemp’s ridley recaptured in Caribbean. Marine Turtle Newsletter 54:13-14.

Witherington, B.E. and K.A. Bjorndal. 1991. Influences of artificial lighting on the seaward orientation of hatchling loggerhead turtles, Caretta caretta. Biological Conservation 55:139-149.

Witherington, B.E. and K.A. Bjorndal. 1991. Influences of wavelength and intensity on hatchling sea turtle phototaxis: implications for sea-finding behavior. Copeia 1991:1060-1069. Witherington&Bjorndal_Copeia1991.pdf


Bjorndal, K.A. 1990. Digestibility of the sponge Chondrilla nucula in the green turtle, Chelonia mydas. Bulletin of Marine Science 47:567-570. Bjorndal_1990_DigestibilityOfTheSpongeChondrillaNucula.pdf

Bjorndal, K.A. and A.B. Bolten. 1990. Goals for sea turtle research in the Gulf of Mexico with respect to the oil and gas industries. Pages 47-58 in Sea turtles and marine mammals of the Gulf of Mexico. Minerals Management Service, U.S. Department of Interior, New Orleans.

Bjorndal, K.A. and A.B. Bolten. 1990. Digestive processing in a herbivorous freshwater turtle: consequences of small intestine fermentation. Physiological Zoology 63:1232-1247.

Bjorndal, K.A., A.B. Bolten, and J.E. Moore. 1990. Digestive fermentation in herbivores: effect of food particle size. Physiological Zoology 63:710-721.

Bolten, A.B. and H.R. Martins. 1990. Kemp’s ridley captured in the Azores. Marine Turtle Newsletter 48:23.

Bolten, A.B., H.R. Martins, M.L. Natali, J.C. Thomé and M.A. Marcovaldi. 1990. Loggerhead released in Brazil recaptured in Azores. Marine Turtle Newsletter 48:24-25.

Magnuson, J.J., K.A. Bjorndal, W.D. DuPaul, G.L. Graham, D.W. Owens, C.H. Peterson, P.C.H. Pritchard, J.I. Richardson, G.E. Saul and C.W. West. 1990. Decline of the sea turtles: causes and prevention. National Academy Press, Washington, D.C. 259 pages.

Moskovits, D.K. and K.A. Bjorndal. 1990. Diet and food preferences of the tortoises Geochelone carbonaria and G. denticulata in northwestern Brazil. Herpetologica 46:207-218.

Norton, T.M., Jacobson, E.R., Sundberg, J.P. 1990. Cutaneous Fibropapilloma an Renal Myxofibroma in a Green Turtle, Chelonia mydas. J. Wildlife Dis. 26:265-270.

Witherington, B.E., K.A. Bjorndal, and C.M. McCabe. 1990. Temporal pattern of nocturnal emergence of loggerhead turtle hatchlings from natural nests. Copeia 1990:1165-1168. Witherington_et_al_Copeia1990.pdf


Bjorndal, K.A. 1989. Flexibility of digestive responses in two generalist herbivores, the tortoises Geochelone carbonaria and Geochelone denticulata. Oecologia 78:317-321.

Bjorndal, K.A. and A.B. Bolten. 1989. Comparison of straight-line and over-the-curve measurements for growth rates of green turtles, Chelonia mydas. Bulletin of Marine Science 45:189-192. BjorndalEtAl_1989_ComparisonOfMeasurementsForGrowthRatesOfGreenTurtles.pdf

Bjorndal, K.A. and A. Carr. 1989. Variation in clutch size and egg size in the green turtle nesting population at Tortuguero, Costa Rica. Herpetologica 45:181-189.

Jacobson, E.R., Mansell, J.L., Sundberg, J. P., Hajjar, L., Reichmann, M.E., Ehrhart, L.M., Walsh, M., Murru, F. 1989. Cutaneous Fibropapillomas of Green Turtles. J. Comp. Path. 101:39-52.

Ogren, L., F. Berry, K. Bjorndal, H. Kumpf, R. Mast, G. Medina, H. Reichart, and R. Witham (editors). 1989. Proceedings of the SecondWestern Atlantic Turtle Symposium (WATS II). NOAA Technical Memorandum NMFS-SEFC-226. 401 pages.


Bjorndal, K.A. and A.B. Bolten. 1988. Growth rates of immature green turtles, Chelonia mydas, on feeding grounds in the southern Bahamas. Copeia 1988:555-564. Bjorndal&Bolten_Copeia1988.pdf

Bjorndal, K.A. and A.B. Bolten. 1988. Growth rates of juvenile loggerheads, Caretta caretta, in the southern Bahamas. Journal of Herpetology 22:480-482. Bjorndal&Bolten_JHerpetol1988.pdf


Bjorndal, K.A. 1987. Digestive efficiency in a temperate herbivorous reptile Gopherus polyphemus. Copeia 1987:714-720. Bjorndal_Copeia_1987.pdf


Bjorndal, K.A. 1986. Effect of solitary versus group feeding on intake in Pseudemys nelsoni. Copeia 1986:234‑235. Bjorndal_Copeia_1986.pdf

Jacobson, E.R., Gaskin, J.M., Roelke, M., Greiner, E.C., Allen, J. 1986. Conjunctivitis, Tracheitis, and Pneumonia Associated with Herpesvirus Infection in Green Sea Turtles. J.A.V.M.A. 189:1020-1023.


Bjorndal, K.A. 1985. Nutritional ecology of sea turtles. Copeia 1985:736‑751. Bjorndal_Copeia_1985_high_res.pdf

Bjorndal, K.A. 1985. Use of ash as an indigestible dietary marker. Bulletin of Marine Science 36:224‑230.

Bjorndal, K.A., A. Carr, A.B. Meylan, and J.A. Mortimer. 1985. Reproductive biology of the hawksbill, Eretmochelys imbricata, atTortuguero, Costa Rica, with notes on the ecology of the species in the Caribbean. Biological Conservation 34:353-368.


Bacon, P., F. Berry, K. Bjorndal, H. Hirth, L. Ogren, and M. Weber (editors). 1984. Proceedings of the Western Atlantic Turtle Symposium. 3 volumes. RSMAS Printing, Miami, Florida.

Bjorndal, K.A. 1984. Nutrition of seagrass herbivores. In: Thayer, G.W., K.A. Bjorndal, J.C. Ogden, S.L. Williams, and J.C. Zieman. Role of larger herbivores in seagrass communities. Estuaries 7:351‑376.


Bjorndal, K.A. and G.H. Balazs (editors). 1983. Manual of sea turtle research and conservation techniques. Center for Environmental Education,Washington, D.C. 126 pages.

Bjorndal, K.A., A.B. Meylan, and B.J. Turner. 1983. Sea turtles nesting at Melbourne Beach, Florida, 1. Size, growth and reproductive biology. Biological Conservation 26:65‑77

Meylan, A.B., K.A. Bjorndal, and B.J. Turner. 1983. Sea turtles nesting at Melbourne Beach, Florida, 2. Post‑nesting movements of Caretta caretta. Biological Conservation 26:79‑90.


Bjorndal, K.A. (editor). 1982. Biology and conservation of sea turtles. Smithsonian Institution Press, Washington, D.C. 583 pages.

Bjorndal, K.A. 1982. The consequences of herbivory for the life history pattern of the Caribbean green turtle. Pages 111-116 in K.A. Bjorndal (ed.), Biology and conservation of sea turtles. Smithsonian Institution Press, Washington, D.C.

Bjorndal, K.A. 1982. Does turning green turtles on their backs affect subsequent reproductive performance? Marine Turtle Newsletter 22:15-16.

Carr, A., A. Meylan, J. Mortimer, K. Bjorndal, and T. Carr. 1982. Surveys of sea turtle populations and habitats in the western Atlantic. NOAA Technical Memorandum NMFS-SEFC-91, 91 pages.

Thayer, G.W., D.W. Engel, and K.A. Bjorndal. 1982. Evidence for short‑circuiting of the detritus cycle of seagrass beds by the green turtle,Chelonia mydas. Journal of Experimental Marine Biology and Ecology 62:173‑183.


Bickham, J.W., K.A. Bjorndal, M.W. Haiduk, and W.E. Rainey. 1980. Karyotype and chromosomal banding patterns of the green turtle (Chelonia mydas). Copeia 1980:540‑543. Bickham_et_al_Copeia_1980.pdf

Bjorndal, K.A. 1980. Nutrition and grazing behavior of the green turtle, Chelonia mydas. Marine Biology 56:147‑154. Bjorndal_1980_NutritionAndGrazingBehaviorOfTheGreenTurtle.pdf

Bjorndal, K.A. 1980. Demography of the breeding population of the green turtle, Chelonia mydas, at Tortuguero, Costa Rica. Copeia 1980:525‑530. Bjorndal_Copeia_1980.pdf

Greiner, E.C., Forrester, D.J. and Jacobson, E.R. 1980. Helminths of Mariculture-reared Green Turtles (Chelonia mydas mydas) from Grand Cayman, B.W.I. Proceedings of the Helminthological Society of Washington, D.C., January 47:142-144


Bjorndal, K.A. 1979. Cellulose digestion and volatile fatty acid production in the green turtle, Chelonia mydas. Comparative Biochemistry and Physiology 63A:127-133. Bjorndal_1979_CelluloseDigestionAndVolatileFattyAcidProductionInTheGreenTurtle.pdf

Bjorndal, K.A. 1979. Urine concentrations of ammonia, urea and uric acid in the green turtle, Chelonia mydas. Comparative Biochemistry and Physiology 63A:509-510.

Carr, A. and K.A. Bjorndal. 1979. Marine turtles of the Bahamas. The Bahamas Naturalist 4:17-20.

Jacobson, E.R., Gaskin, J.M., Shields, R.P. and White, F.H. 1979. Mycotic Pneumonia of Mariculture Reared Green Sea Turtles. J.A.V.M.A. 175:929-933.


Bjorndal, K.A. and A.B. Bolten. 1978. Union Creek–a look to the future. The Bahamas Naturalist 4:18‑27.


Bjorndal, K.A. 1977. Turtle grass, Thalassia testudinum. The Bahamas Naturalist 2:17-19.




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