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Dr. Sadie J. Ryan

sadie-research-areas   RYAN_150_200

Dr. Sadie J. Ryan

Associate Professor, Medical Geography

Ryan Lab website

Focus Areas

Areas of Specialization

  • Medical Geography
  • Disease Ecology
  • Tropical conservation
  • Human-wildlife interface
  • Spatial models

Educational Background

  • PhD in Environmental Science, Policy and Management (ESPM), University of California at Berkeley, 2006
  • B.A. in Ecology and Evolutionary Biology (EEB), Princeton University, 1998

Recent Courses

  • People and Plagues (GEO 3930)

Current Graduate Students


Postdoctoral Researchers

  • Phillip Boersch-Supan, PhD, co-mentor with Dr. L. Johnson, Department of Integrative Biology, University of South Florida “Quantifying how Bioenergetics and Foraging Determine Population Dynamics in Threatened Antarctic Albatrosses”. Funded by NSF

Undergraduate Researchers

  • Taylor Rouviere
  • Stephen MacKay
  • Katie Walker

Recent Graduate Students

  • Emily Gavard (M.S. 2017, SUNY-ESF)
  • Tess Youker (M.S. 2016, SUNY-ESF)
  • Rebecca Fuda (M.S., 2015, SUNY-ESF)
  • Andrea Thomen (M.S., 2014, SUNY-ESF)
  • Lindsey Scales (M.P.S., 2014, SUNY-ESF)
  • Dr. Luis Escobar, (Postdoc, 2013-2014, SUNY-UMU)

In My Own Words

I focus on medical geography and ecology at the human-wildlife interface, particularly where it pertains to disease ecology, conservation biology, and wildlife management. I use methods from landscape ecology (GIS, remote sensing, spatial analyses), quantitative ecology, and epidemiology to analyze landscape change, health outcomes, and the impact of anthropogenic changes, including climate change.

Recent Publications

An up to date list of Dr. Ryan’s publications is available on the Ryan Lab Website.

(PDFs are intended for single copy, research use only, to comply with copyright regulations)

(completed manuscripts, posted to an archive, many in review somewhere)
Ryan, S.J. et al. 2017. Climate change drives uncertain global shifts in potential distribution and seasonal risk of Aedes-transmitted viruses.
BioRXiv doi: 

Kenneson, A., et al. 2017. Social-ecological factors and preventive actions decrease the risk of dengue infection at the household-level: results from a prospective dengue surveillance study in Machala, Ecuador. BioRXiv doi

Carlson, C. et al. 2017. Consensus and conflict among ecological forecasts of Zika virus outbreaks in the United States. BioRXiv doi

James, TG and Ryan, S.J. 2017. Differences in STI knowledge accuracy and STI/HIV testing among a random sample of college students: A secondary survey analysis. SocArXiv doi

Nightingale et al., 2017. Dengue fever and Aedes aegypti risk in the Galápagos Islands, Ecuador. BioRXiv doi:

Lippi et al., 2017. The social and spatial ecology of dengue presence and burden during an outbreak in Guayaquil, Ecuador, 2012. 
BioRXiv doi:

Ryan, S.J. et al. 2017.​ Environmental cholera (Vibrio cholerae) dynamics in an estuarine system in southern coastal Ecuador.
​​BioRXiv doi:

Stewart-Ibarra, A.M., et al. 2017. The high burden of dengue and chikungunya in southern coastal Ecuador: Epidemiology, clinical presentation, and phylogenetics from a prospective study in Machala in 2014 and 2015. BioRXiv doi:

Ryan, S.J., Carlson, C., Stewart-Ibarra, A.M., Borbor-Cordova, M.J., Romero, M.M., Cox, S., Mahon, R., Trotman, A., St Ville, E.S., Ahmed, S. accepted. Zika Virus Outbreak, Dominica, 2016. Emerging Infectious Diseases

Bardosh, K.L., Jean, L., Beau de Rochers, V.M., Lemoine, J.F., Okech, B., Ryan, S.J., Welburn, S., Morris, J. G. 2017Polisye Kont Moustik: A culturally competent approach to larval source reduction in the context of lyphatic filariasis and malaria elimination in Haiti. Tropical Medicine and Infectious Disease.​ 2(39) [ PDF]

Johnson, L.R., Gramacy, R.B., Cohen, J., Mordecai, E., Murdock, C., Rohr, J., Ryan, S.J., Stewart-Ibarra, A., Weikel, D. accepted. Phenomenological forecasting of disease incidence using heteroskedastic Gaussian processes: a dengue case study. Annals of Applied Statistics. Preprint: ArXiv

Johnson, L.R., Boersch-Supan, P.H., Phillips, R.A., Ryan, S.J. accepted. Changing measurements or changing movements? Sampling scale and movement model identifiability across generations of biologging technology. Ecology and Evolution. Preprint: ArXiv:

MacKenzie, C.A., Fuda, R.K., Ryan, S.J., Hartter, J. in press. Drilling through Conservation Policy: Oil Exploration in Murchison Falls Protected Area, Uganda. Conservation & Society

Dobbins, M., Steinberg, M., Broadbent, E., Ryan, S.J. accepted. Habitat use, activity patterns, and human interactions with jaguars in southern Belize. Oryx.

Pettorelli, N., Nagendra, H., Rocchini, D., Rowcliffe, M., Williams, R., Ahumada, J., De Angelo, C., Atzberger, C., Boyd, D., Buchanan, G., Chauvenet, A., Disney, M., Duncan, C., Fatoyinbo, T., Fernandez, N., Haklay, M., He, K., Horning, N., Kelly, N., de Klerk, H., Liu, X., Merchant, N., Paruelo, J., Roy, H., Roy, S., Ryan, S., Sollmann, R., Swenson, J. and Wegmann, M. 2017. Remote Sensing in Ecology and Conservation: three years on. Remote Sens Ecol Conserv, 3: 53–56. [ PDF]

Mordecai, E.A., Cohen, J., Evans, M.V., Gudapati, P., Johnson, L.R., Lippi, C.A., Miazgowicz, K., Murdock, C.C., Rohr, J.R., Ryan, S.J., Savage, V., Shocket, M., Stewart Ibarra, A.M., Thomas, M.B., Weikel, D.P. 2017. Detecting the impact of temperature on transmission of Zika, dengue, and chikungunya using mechanistic models. PLOS NTDs. 11(4): e0005568 [ PDF]

Ryan, S.J., Palace, M., Hartter, J., Diem, J.E., Chapman, C.A., Southworth, J. 2017Population pressure and global markets drive a decade of forest cover change in Africa’s Albertine Rift. Applied Geography. 81:52-59 [ PDF]

Southworth J., Ryan S.J., Bunting E., Herrero H.V., Nagendra H., Gibbes C., Agarwal S. 2017. Protected Areas, Climate Change, and Ecosystem Sustainability, Reference Module in Earth Systems and Environmental Sciences, Elsevier, 2017. [ PDF available by request from authors, subject to terms of Elsevier publishing]

Boersch-Supan, P., Ryan, S.J., Johnson, L.R. 2017. deBInfer: Bayesian inference for dynamical models of biological systems in R. Methods in Ecology and Evolution 8(4): 511-518. [ PDF] (R package on CRAN)

Youker-Smith, T.E., Whipps, C.M., Ryan, S.J. 2016. Detection of an FV3-like Ranavirus in Wood Frogs (Lithobates sylvaticus) and Green Frogs (Lithobates clamitans) in a Constructed Vernal Pool Network in Central New York State. Herpetological Review. 47(4): 595-598 [ PDF]

Fuda, R.K., Ryan, S.J., Cohen, J., Hartter, J., Frair, J. 2016. Assessing impacts to primary productivity at the park edge in Murchison Falls Conservation Area, Uganda. Ecosphere. 7(10):e01486. 10.1002/ecs2.1486 [ PDF]

Ryan, S.J., Gavard, E.J., Cheeseman, A.E., Cohen, J.B., Whipps, C.W. 2016. Reference and baseline hematocrit measures for the threatened New England cottontail (Sylvilagus transitionalis) and comparison with sympatric Eastern cottontail (Sylvilagus floridanus) rabbits. Journal of Zoo and Wildlife Medicine. 47(2):659-662 [ PDF]

Cameron, E.Z.* and Ryan, S.J.* 2016Welfare at multiple scales: importance of zoo elephant population welfare in a world of declining wild populations. Invited formal comment. PLOS ONE 11(7): e0158701 [ PDF]
Formal comment on the co-edited collection Epidemiological Investigations of North American Zoo Elephant Welfare (

Handel A., Beltran, E., Borbor Cordova M.J., Fessler A., Finkelstein J., Robalino, R., Ryan S.J., Stewart Ibarra A.M. 2016 Knowledge, attitudes, and practices regarding dengue infection among healthcare providers in Machala, Ecuador. BMC Tropical Diseases, Travel Medicine and Vaccines. (2):8 Online:  DOI: 10.1186/s40794-016-0024-y [ PDF]

Borbor-Cordova, M., Ayala, E., Cardenas, W., Endy, T., Finkelstein, J., Leon, R., Muñoz, Á., Mejía, R., Polhemus, M., Recalde Coronel, G., Ryan, S., Stewart-Ibarra, A. 2016. “Chapter 5: Developing and delivering health-tailored climate products and services, Case study 5.C Vector-virus microclimate surveillance system for dengue control in Machala, Ecuador”, Climate Services for Health: Global case studies of enhancing decision support for climate risk management and adaptation, WHO/WMO, Geneva. pp.106-109 [ PDF]

Taylor, R., Ryan, S.J., Brashares, J., Johnson, L.R. 2016Hunting, food subsidies, and mesopredator release: the dynamics of crop-raiding baboons in a managed landscape. Ecology 97(4): 951-960 [ PDF] (Supporting Data Publication: Ryan, S. J2015. Primate life history data.

Hartter, J., Dowhaniuk, N., MacKenzie, C.A., Ryan, S.J., Diem, J.E., Palace, M.W., Chapman, C.A. 2016Perceptions of Risk in Communities near Parks in an African Biodiversity Hotspot. Ambio. 45(6), 692-705  ​[ PDF]

Southworth, J., Zhu, L., Bunting, E., Ryan, S.J., Herrero, H., Waylen, P.R., Hill, M.J. 2016Changes in Vegetation Persistence across Global Savanna Landscapes, 1982-2010. Journal of Land Use Science, 11(1): 7-32 . [ PDF]

Data Publication: Ryan, S. J2015. Primate life history data.

Ryan, S.J., McNally, A., Johnson, L.R., Mordecai, E.A., Ben-Horin, T., Paaijmans, K.P., Lafferty, K.D. 2015. Mapping physiological suitability limits of malaria in Africa under climate change. Journal of vector borne and zoonotic diseases 15(12): 817-725 [ PDF]

Ryan, S.J., Ben-Horin, T., Johnson, L.R. 2015. Malaria control and senescence: the importance of accounting for the pace and shape of ageing in wild mosquitoes. Ecosphere 6(9):170 [ PDF]

Escobar, L.E., Ryan, S.J., Stewart-Ibarra, A.M., Finkelstein, J.L., King, C.A., Qiao, H., Polhemus, M.E. 2015A Global Map of Suitability for Coastal Vibrio cholerae Under Current and Future Climate Conditions. Acta Tropica 149: 202-211 [ PDF]

Ryan, S.J., Southworth, J., Hartter, J., Dowhaniuk, N., Fuda, R., Diem, J. 2015Household level influences on fragmentation in an African park landscape. Applied Geography 58: 18-31[ PDF]

Johnson, L.R.  Ben-Horin, T., Lafferty, K.D., McNally, A., Mordecai, E., Paaijmans, K.P., Pawar, S., Ryan, S.J. 2015. Understanding uncertainty in temperature effects on vector-borne disease: A Bayesian approach. Ecology 96(1): 203-213. [ PDF]

Hartter, J., Ryan, S.J., Mackenzie, C.A., Goldman, A., Dowhaniuk, N., Palace, M.W., Diem, J.E., Chapman, C.A. 2015Now there is no land: A story of ethnic migration in a protected area landscape in western Uganda. Population and Environment 36(4): 452-479 [ PDF]

Stewart Ibarra, A.M., Munoz, A.G., Ryan, S.J., Borbor, M.J., Beltran Ayala, E., Finkelstein, J.L., Mejia, R., Ordonez, T., Recalde Coronel, G.C., Rivero, K. 2014Spatiotemporal clustering, climate periodicity, and social-ecological risk factors for dengue during an outbreak in Machala, Ecuador, in 2010. BMC Infectious Diseases 14:610 [ PDF]

L.R. Johnson, K.D. Lafferty, A. McNally, E. Mordecai, K. Paaijmans, S. Pawar, S.J. Ryan. 2014. “Mapping the distribution of Malaria: Current Approaches and Future Directions”. in Analyzing and Modeling Spatial and Temporal Dynamics of Infectious Diseases. D. Chen, B. Moulin, J. Wu (Eds). John Wiley & Sons. pp 189-209 [ PDF] [ BOOK LINK ]

Stewart Ibarra, A.M., Luzadis, V.A., Borbor Cordova, M.J., Silva, M., Ordoñez, T., Beltrán Ayala, E., Ryan, S.J. 2014A social-ecological analysis of community perceptions of dengue fever and Aedes aegypti in Machala, Ecuador. BMC Public Health 14:1135 [ PDF]

Diem, J.E., Hartter, J., Ryan, S.J., Palace, M.W. 2014. Validation of satellite rainfall products for western Uganda. Journal of Hydrometeorology. 15(5): 2030-2038. [ PDF]

Cornélis, D., Melletti, M., Korte, L., Ryan, S.J., Mirabile, M., Prin, T., and Prins, H.H.T. 2014. Species Account: African buffalo (Syncerus caffer Sparrman, 1779). in
Ecology, Evolution and Behaviour of Wild Cattle: Implications for Conservation. Melleti, M. and Burton, J., (Eds). Cambridge. pp 326-372 [ PDF] [ BOOK LINK]

Diem, J.E., Ryan, S.J., Hartter, J., Palace, M.W. 2014. Satellite-based rainfall data reveal a recent drying trend in central equatorial Africa. Climatic Change. 126(1-2): 263-272. [ PDF]

Hartter, J., Solomon, J., Ryan, S.J., Jacobson, S.K., Goldman, A. 2014. Contrasting perceptions of ecosystem services of an African forest park. Environmental Conservation 41(4): 330-340. [ PDF]

Dowhaniuk, N., Hartter, J., Ryan, S.J. 2014Impact of spatial data discrepancies on protected areas management: an example from East Africa. Environmental Management 54(3): 596-605. [ PDF]

Stewart Ibarra, A.M., Ryan, S.J., Beltran, E. Mejía, R., Silva, M., Muñoz, A. 2013Dengue vector dynamics (Aedes aegypti) influenced by climatic and social factors in Ecuador: implications for targeted control. PLOS ONE 8(11): e78263 [ PDF]

Ryan, S.J., Jones, J.H., Dobson, A.P. 2013Interactions between social structure, demography, and transmission determine disease persistence in primates. PLOS ONE 8(10): e7686 [ PDF]
Supplementary Discussion S1: From theory to reality: how can we establish and understand contact rates for these transmission modes? [ PDF]

Hartter, J.*, Ryan, S.J.*, MacKenzie, C.A., Parker, J.N., Strasser, C. Spatially explicit data: stewardship and ethical challenges in science. PLOS Biology 11(9): e1001364 [ PDF]

Mordecai, E.A., Paaijmans, K.P., Johnson, L.R., Balzer, C.H., Ben-Horin, T., deMoor, E., McNally, A., Pawar, S., Ryan, S.J., Smith, T.C., Lafferty, K.D., 2013. Physiological constraints dramatically lower the expected temperature for peak malaria transmission. Ecology Letters 16(1):22-30. [ PDF]

Ryan, S.J. and Hartter, J. 2012. Beyond ecological success of corridors: integrating land use history and demographic change to provide a whole landscape perspective. Ecological Restoration. 30(4): 320-328 [ PDF] Invited paper: Special Issue on Design Approaches to Ecological Restoration in Ecological Restoration.

M.J. Tildesley and Ryan, S.J. 2012. Disease prevention versus data privacy:using landcover maps to inform spatial epidemic models. PLOS Computational Biology8(11): e1002723 [ PDF]

Rudnick, D., Ryan, S.J., Preziosi, D., Epps, C., Deiffenbach, F., Kintsch, J., Beier, P., Jenness, J., Perkl., R., Trombulak, S., Hartter, J., Gerber, L., Merenlender, A., Austen, D., Cushman, S. 2012. The role of landscape connectivity in planning and implementing conservation and restoration priorities. Issues in Ecology 16. [ PDF]

Ryan, S.J., Cross, P.C., Winnie, J, Jr, Hay, C., Bowers, J., Getz, W.M. 2012. The utility of NDVI for predicting African buffalo forage quality. Journal of Wildlife Management. 76(7):1499-1508  [ PDF] [ COVER]
Data files: DOI: 10.5061/dryad.7jd83    
Fecal Nitrogen and Phosphorous measurements: NP_01-02_KNP
Buffalo Body Condition Data
Vegetation and fecal indicators of Nitrogen and Phosphorous

Ryan, S.J., Brashares, J. S, Walsh, C., Milbers, K., Kilroy, C., Chapman, C.A. 2012. A survey of gastrointestinal parasites of olive baboons (Papio anubis) in human settlement areas of Mole National Park, Ghana. J. Parasitology 98:885-888. [ PDF]

Hartter, J., Stampone, M.D., Ryan, S.J., Kirner, K., Chapman, C.A., Goldman, A., 2012. Patterns and perceptions of climate change around a biodiversity hotspot. PLOS ONE 7(2): e32408 [ PDF]

Ryan, S.J. and Walsh, P.D. 2011. Consequences of non-intervention for infectious disease in African great apes. PLOS ONE 6(12): e29030. [ PDF]

Wolf, A., Anderegg, W., Ryan, S.J., Christensen, J. 2011. Robust detection of plant species range shifts under biased sampling regimes. Ecosphere 2(10):115 [ PDF]

Hartter, J.* and Ryan, S.J.*, Southworth, J., and Chapman, C.A. 2011. Landscapes as continuous entities: forest disturbance and recovery in the Albertine Rift Landscape. Landscape Ecology 26(6):877-890 [ PDF]

Stampone, M., Hartter, J., Chapman, C.A., Ryan, S.J. 2011.Localized precipitation trends in and around a forest park in east equatorial Africa. Research Journal of Environmental and Earth Sciences 3(1):14-23 [PDF]

Petorelli, N., Ryan, S.J., Mueller, T., Bunnefeld, N., Jedrzejewska, B., Lima, M., Kausrud, K. 2011. The Normalized Difference Vegetation Index (NDVI): unforeseen successes in animal ecology. Climate Research 46(1):15-27 [PDF]

2006 – 2010
Hartter, J. and Ryan, S.J. 2010. Top-down or bottom-up? Decentralization, natural resource management and usufruct rights in the forests and wetlands of Western Uganda. Land Use Policy 27(3): 815-826[ PDF]

Ladau, J. and Ryan, S.J. 2010. MPowering ecologists: community assembly tools for community assembly rules. Oikos 119(7):1064-1069 [ PDF   ]

Chapman, C.A., Huffman,M.A., Ryan,S.J., Sengupta, R., and  Goldberg, T. 2009. “Ways forward in the study of primate parasite ecology”. in Primate parasite ecology: the dynamics and study of host-parasite relationships. Huffman, M.A. and Chapman, C.A., eds. Cambridge Studies in Biological and Evolutionary Anthropology. Cambridge University Press. [PDF]

Sánchez, M.S., Lloyd-Smith, J.O., Williams, B.G., Porco, T.C., Ryan, S.J., Borgdorff, M.W., Mansoer, J., Dye, C. and Getz, W.M. 2009. Incongruent HIV and tuberculosis co-dynamics in Kenya: Interacting epidemics monitor each other. Epidemics 1(1):14-20 [ PDF]

Bar-David, S, Bar-David, I, Cross, P.C. Ryan, S.J., Knechtel, C.U., Getz, W.M. 2009. Methods for assessing movement path recursion with application to African buffalo in South Africa. Ecology 90(9):2467–2479 [PDF]

Ryan, S.J., Starks, P.T., Milton, K. and Getz, W.M. 2008. Intersexual conflict and group size in Alouatta palliata: a 23-year evaluation.  International Journal of Primatology. 29:405-420  [PDF]

Gusset, M.* and Ryan, S.J.*, Hofmeyr, M., Van Dyk, G., Davies-Mostert, H.T., Graf, J.A., Owen, C., Szykman, M., Macdonald, D.W., Monfort, S.L., Wildt, D.E., Maddock, A.H., Mills, M.G.L., Slotow, R., Somers, M.J. 2008. Efforts going to the dogs? Evaluating attempts to re-introduce endangered wild dogs in South Africa. Journal of Applied Ecology 45:100-108  [PDF]

Ryan, S.J., Knechtel, C.U. and Getz, W.M. 2007. Ecological cues, gestation length and birth timing in African Buffalo (Syncerus caffer). Behavioral Ecology 18:635-644 [ PDF

Getz, W.M., Fortmann-Roe, S, Cross, P.C., Lyons, A.J. Ryan, S.J., et al. 2007. LoCoH: Nonparametric kernel methods for constructing home ranges and utilization distributions. PLOS ONE 2(2): e207. [ PDF]

Ryan, S.J. 2006. The role of culture in conservation of small or endangered populations. Journal of Conservation Biology 20(4):1321-1324 [PDF]

Ryan, S.J., Knechtel, C.U. and Getz, W.M.2006. Range and habitat selection of African buffalo in South Africa. Journal of Wildlife Management 70(3):764-776 [PDF]

Redfern, J.V., Ryan, S.J. and Getz, W.M. 2006. Defining Herbivore Assemblages in the Kruger National Park: A Correlative Coherence Approach. Oecologia 146:632-640 [PDF]

Ryan, S.J., and Getz, W.M. 2005. A spatial location allocation GIS framework for managing water sources in a savanna nature reserve. South African Journal of Wildlife Research 35(2):153-178 [PDF]

Ryan, S.J. and Jordaan, W. 2005. Activity Patterns of African Buffalo Syncerus caffer in the Lower Sabie Region, Kruger National Park, South Africa. Koedoe 48(2): 117-124 [PDF]

Faust, L.J., Thompson, S.D., Earnhardt, J.M., Brown, E., Ryan, S.J., Sherman, M. and Yurenka, M. 2003. Using Stage-Based System Dynamics Modeling for Demographic Management of Captive Populations. Zoo Biology, 22:1, Wiley-Liss [PDF]

Ryan, S., Roth, A.M., Thompson,S.D. and Gold, K.C. 2002. Effects of hand rearing on the reproductive success of Western Lowland Gorillas in North America. Zoo Biology 21:3, Wiley-Liss [PDF]

Ryan, S.J. and Thompson, S.D., 2001. Disease Risk and Inter-institutional Transfer of Specimens in Cooperative Breeding Programs: Herpes and the Elephant Species Survival Plans, Zoo Biology 20:2, Wiley-Liss [PDF]

Dobson, A., Ralls. K., Foster, M., Soule, M.,Simberloff, D., Doak, D., Estes, J., Mills, S., Mattson, D., Dirzo, R., Arita, H., Ryan, S., Norse, E., Noss, R., Johns, D. 1999. “Corridors: Reconnecting Fragmented Landscapes”. In Continental Conservation, Michael E. Soule and John Terborgh, eds. The Wildlands Project,Island Press. [PDF]


Ryan, S. J., Rheingans, R., Amratia, P., Amaya, M.P., Laytner, L. A., Bagamian, K. H., Anderson, J.D., Watson, J., McNamara, K., Cumming, O. 2016. “WASH Poverty Diagnostic; Poverty Risk Model Assessment: Ecuador.” World Bank Report, 51pp. Report commissioned by the “Poverty Risk Models (PRM) for water, sanitation and health (WASH) project” for World Bank, Washington, D.C.

Rheingans, R., Bagamian, K. H., Anderson, J. D., Ryan, S. J., Amratia, P., Amaya, M. P., Bouland, J., Laytner, L. A., Watson, J., Cumming, O. 2016. “WASH Poverty Diagnostic; Poverty Risk Model Assessment: Bangladesh.” World Bank Report, 56pp. Report commissioned by the “Poverty Risk Models (PRM) for water, sanitation and health (WASH) project” for World Bank, Washington, D.C.

Rheingans, R., Anderson, J. D., Bagamian, K. H., Ryan, S. J., McNamara, K., Laytner, L. A., Amratia, P., Watson, J., Cumming, O. 2016. “WASH Poverty Diagnostic; Poverty Risk Model Assessment: Democratic Republic of Congo.” World Bank Report, 56pp. Report commissioned by the “Poverty Risk Models (PRM) for water, sanitation and health (WASH) project” for World Bank, Washington, D.C.

Rheingans, R., Bagamian, K. H., Anderson, J. D., Ryan, S. J., Laytner, L. A., McNamara, K., Watson, J., Cumming, O. 2016. “WASH Poverty Diagnostic; Poverty Risk Model Assessment: Ethiopia.” World Bank Report, 56pp. Report commissioned by the “Poverty Risk Models (PRM) for water, sanitation and health (WASH) project” for World Bank, Washington, D.C.

Rheingans, R., Bagamian, K. H., Anderson, J. D., Ryan, S. J., Laytner, L. A., McNamara, K., Amaya, M. P., Watson, J., Cumming, O. 2016. “WASH Poverty Diagnostic; Poverty Risk Model Assessment: Haiti.” World Bank Report, 54pp. Report commissioned by the “Poverty Risk Models (PRM) for water, sanitation and health (WASH) project” for World Bank, Washington, D.C.

Rheingans, R., Anderson, J.D., Bagamian, K.H., Ryan, S.J., Watson, J., Laytner, L.A., Cumming, O. 2016. “WASH Poverty Diagnostic; Poverty Risk Model Assessment: Mozambique.” World Bank Report, 62pp. Report commissioned by the “Poverty Risk Models (PRM) for water, sanitation and health (WASH) project” for World Bank, Washington, D.C.

Rheingans, R., Bagamian, K. H., Anderson, J.D., Ryan, S. J., Laytner, L. A., McNamara, K., Watson, J., Cumming, O. 2016. “WASH Poverty Diagnostic; Poverty Risk Model Assessment: Nigeria.” World Bank Report, 54pp. Report commissioned by the “Poverty Risk Models (PRM) for water, sanitation and health (WASH) project” for World Bank, Washington, D.C.

Rheingans, R., Anderson, J. D., Bagamian, K. H., Ryan, S.J., Amaya, M. P., Laytner, L. A., McNamara, K., Watson, J., Cumming, O. 2016. “WASH Poverty Diagnostic; Poverty Risk Model Assessment: Tajikistan.” World Bank Report, 54pp. Report commissioned by the “Poverty Risk Models (PRM) for water, sanitation and health (WASH) project” for World Bank, Washington, D.C.

Rheingans, R., Bagamian, K. H., Anderson, J. D., Ryan, S. J., Watson, J., Amratia, P., Laytner, L. A., Cumming, O. 2016. “WASH Poverty Diagnostic; Poverty Risk Model Assessment: Pakistan.” World Bank Report, 58pp. Report commissioned by the “Poverty Risk Models (PRM) for water, sanitation and health (WASH) project” for World Bank, Washington, D.C.

Bardosh, K, Ebi, K., Ryan, S.J., Welburn, S., Singer. B, 2015. “Addressing Vulnerability, Building Resilience: Community-based Adaptation to Vector-Borne Diseases in the Context of Global Change.” Review commissioned by the UNICEF/UNDP/World Bank/WHO Special Programme for Research and Training in Tropical Diseases (TDR) as part of the research initiative “Population Health Vulnerabilities to Vector-Borne Diseases: Increasing Resilience under Climate Change Conditions in Africa.”With funding support from the International Development Research Centre (IDRC) of Canada.

Ryan, S.J., 2011. Invited Book Review. “Governing Africa’s Forests in a Globalized World Laura A German, Alain Karsenty and Anne-Marie Tiani (Editors)”. Natural Resources Forum 35(2): 146-147[PDF]

Hartter, J., Ryan, S.J., Southworth, J., and Chapman, C.A. 2010. “Fortresses and Fragments: Impacts of Fragmentation in a Forest Park Landscape”. in Proceedings of IUFRO Landscape Ecology International Conference. Braganca, Portugal [PDF]

Getz, W.M, Cross, P.C., Jolles, A.E., Lloyd-Smith, J.O., Ryan, S.J., Baxter, P.W.J., Bowers, J., Hay, C.T., Knechtel, C., Tambling, C.J., Turner, W.C., duToit, J.T. 2005. “Bovine Tuberculosis in the African Buffalo: The Role of Population Models”. in Conservation and development interventions at the wildlife/livestock interface: implications for wildlife, livestock and human health. Osofsky, S. A., S. Cleaveland, et al., Eds. Gland, Switzerland and Cambridge, UK, IUCN

Ryan, S., Roth, A.M., Thompson, S.D., Gold, K.C. 2001. “Effects of hand rearing on the reproductive success of Western Lowland Gorillas in North America”. in The Apes: Challenges for the 21st Century. Chicago, Chicago Zoological Society, p.99. [PDF]

Faust, L. J., Thompson, S.D., Earnhardt, J.E., Sherman, M., Brown, E. and Ryan, S. J. 2001 . “Innovative techniques in population demography to improve captive management of western lowland gorillas”. Brookfield Zoo, 2001. in The Apes: Challenges for the 21st Century. Chicago, Chicago Zoological Society, p.207. [PDF]

Ryan, S.J. 2006. Spatial Ecology of African Buffalo and their Resources in a Savanna Ecosystem. PhD Dissertation, Department of Environmental Science, Policy and Management, UC Berkeley. [ PDF]

Ryan, S.J. 1998. Primate Concerns: Development of a Rapid Censusing Technique, Social Effects on Pathogen Transmission and an Examination of Primate Conservation. BA Thesis, Department of Ecology and Evolutionary Biology, Princeton University

Ryan, S., and Nadolny, R. 2011 “Report on the Second Advanced Study Institute on Mathematical Modeling in Conservation Biology at the Kenya Wildlife Service Training Institute”. Society for Mathematical Biology newsletter, May 2011, p.6-7 [ PDF]

Smith, C, with Ouzman, S., Ryan, S. and Schonfeldt-Aultman, S. 2003. ” Renboogtaal in Renboogdorp gepraat”. Die Volksblad 15 May

Ryan, S.J.(ed.). 1999. North American Regional Studbook for Sunbittern (Eurypyga helias). AZA /Lincoln Park Zoo
Ryan, S.J.(ed.). 1999. North American Regional Studbook for Pygmy Hippos (Hexaprotodon liberiensis). AZA/LPZ
Ryan, S.J.(ed.). 2000. North American Regional Studbook for Snow Leopards (Pantera uncia). AZA /LPZ
Ryan, S.J.(ed.). 2000. AZA Research Coordinators Directory. AZA/LPZ

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